Post on 09-Mar-2016
description
664 FAMILY VI. STREPTOCOCCACEAE
TAB
LE 1
31.
Ch
arac
teri
stic
s of
Str
epto
cocc
us s
peci
esa
Cha
ract
eris
tic
S. pyogenes
S. acidominimus
b
S. agalactiae
S. alactolyticus
S. anginosus
S. australis
S. canis
S. constellatus subsp.
constellatus
S. constellatus subsp.
pharyngis
S. criceti
S. cristatus
S. devriesei
S. didelphis
Cel
l siz
e (m
) 0.
51.
0 N
D
0.6
1.2
0.5
1.0
0.5
1.0
ND
N
D
0.5
1.0
0.5
1.0
0.5
~1.0
N
D
ND
C
atal
ase
+c
Acid
pro
duct
ion
from
:
N-
Ace
tyl-g
luco
sam
ine
+ N
D
+ +
d N
D
+ +
+ +
+
ND
Ado
nito
l
N
D
ND
ND
N
D
ND
N
D
N
D
A
myg
dalin
ND
d +
ND
d +
+
+ N
D
A
rabi
nose
A
rbut
in
N
D
d
+ N
D
+ d
+ +
+ +
ND
Ara
bito
l
N
D
ND
ND
N
D
ND
N
D
e
ND
Cel
lobi
ose
d N
D
d +
d N
D
ND
d
ND
+
ND
+
ND
Cyc
lode
xtrin
ND
N
D
ND
N
D
ND
ND
Dex
tran
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
Dul
cito
l
N
D
ND
ND
N
D
ND
N
D
d N
D
Er
ythr
itol
ND
N
D
N
D
ND
N
D
ND
ND
Escu
lin
ND
N
D
N
D
ND
N
D
ND
N
D
ND
N
D
ND
+
ND
Fruc
tose
+
+ +
+ N
D
ND
+
ND
N
D
ND
N
D
+ N
D
Fu
cose
N
D
N
D
N
D
ND
ND
N
D
ND
N
D
N
D
G
alac
tose
+
ND
+
+ N
D
ND
+
ND
N
D
+ d
+ N
D
G
entio
bios
e N
D
ND
N
D
d N
D
ND
ND
N
D
ND
N
D
+ N
D
G
luco
nate
N
D
ND
ND
N
D
ND
N
D
+ N
D
2-
Ket
oglu
cona
te
ND
ND
ND
N
D
N
D
ND
N
D
ND
ND
5-K
etog
luco
nate
N
D
N
D
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
G
luco
se
+ +
+ +
+ N
D
+ +
+ +
ND
+
ND
Gly
cero
l d
ND
+
d
ND
N
D
N
D
N
D
G
lyco
gen
d
g
N
D
N
D
In
osito
l N
D
N
D
N
D
ND
ND
N
D
ND
N
D
N
D
In
ulin
N
D
+
ND
+
d
La
ctos
e +
+ d
+
+ +
d +
d d
+ d
Ly
xose
N
D
N
D
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
M
alto
se
+ N
D
+ +
+ +
+ +
ND
+
+ +
ND
Man
nose
N
D
ND
N
D
+ N
D
ND
+
ND
N
D
ND
N
D
+ N
D
M
anni
tol
+
dh
()
dh (
)
+
+
Mel
ibio
se
N
D
d
d
+
+
ND
Mel
ezito
se
N
D
d
ND
N
D
ND
ND
Met
hyl-D
-man
nosi
de
ND
N
D
N
D
ND
N
D
ND
ND
Met
hyl-D
-glu
cosi
de
+
+ d
+
d d
ND
ND
+
ND
-Met
hylx
ylos
ide
ND
N
D
ND
N
D
ND
N
D
N
D
ND
N
D
ND
ND
Met
hyl-D
-xyl
osid
e
d
N
D
ND
ND
N
D
ND
N
D
N
D
Pu
llula
n d
+
d
+ N
D
N
D
ND
d
N
D
R
affin
ose
N
D
+ d
+
+ d
R
ham
nose
N
D
N
D
N
D
ND
ND
N
D
N
D
R
ibos
e
+
+
N
D
+
Sa
licin
+
ND
d
d +
ND
+
ND
N
D
+ N
D
+
So
rbito
l
+
+
So
rbos
e
N
D
ND
ND
N
D
d N
D
St
arch
N
D
N
D
d N
D
ND
+
ND
N
D
N
D
ND
d
Su
cros
e +
+ +
+ +
+ +
ND
N
D
+ N
D
+ N
D
Ta
gato
se
N
D
d
ND
N
D
+
ND
Treh
alos
e +
+
d +
d
d +
d +
+ +
D
-Tur
anos
e N
D
N
D
d N
D
ND
ND
N
D
ND
N
D
X
ylos
e
N
D
ND
ND
N
D
N
D
N
D
X
ylito
l N
D
N
D
N
D
ND
ND
N
D
ND
N
D
Hyd
roly
sis o
f:
Arg
inin
e +
+
+
+ +
+ +
dh
(+)
dh
(+)
Es
culin
d
+
+ +
ND
+
+ +
dh (+
)
ND
G
elat
in
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
GENUS I. STREPTOCOCCUS 665
(con
tin
ued)
Hip
pura
te
+h
, m (
) +
N
D
+ +h
()
St
arch
ND
ND
ND
N
D
ND
Prod
uctio
n of
:o
A
cid
phos
phat
ase
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
Alk
alin
e ph
osph
atas
e +
+
+
+ +
+ +
ND
+
-L
-Ara
bino
sida
se
ND
N
D
ND
N
D
N
D
ND
N
D
ND
N
D
ND
N
D
ND
-D-G
alac
tosi
dase
ND
d
+
d
ND
+ d
-D
-Gal
acto
sida
se
N
D
d
p +
+
ND
d
Ph
osph
o--g
alac
tosi
dase
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
-D
-Glu
cosi
dase
N
D
ND
N
D
ND
d
ND
N
D
+ +
ND
ND
N
D
-D
-Glu
cosi
dase
ND
d
+
ND
+ N
D
+
-L-F
ucos
idas
e N
D
ND
N
D
ND
ND
N
D
ND
+
ND
N
D
-D
-Fuc
osid
ase
ND
N
D
ND
N
D
N
D
ND
+ N
D
N
D
ND
-L-F
ucos
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
-D
-Glu
curo
nida
se
d +
d
d
N
D
ND
+
-Glu
cosa
min
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
N-
Ace
tyl-
-D-g
alac
tosa
min
idas
e N
D
ND
N
D
ND
ND
N
D
+
ND
+
ND
N
D
N-
Ace
tyl-
-D-g
luco
sam
inid
ase
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N-A
cety
l--D
-glu
cosa
min
idas
e
p
ND
d N
D
+ N
D
ND
-Mal
tosi
dase
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
-M
alto
sida
se
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
Gly
cyl-t
rypt
opha
n ar
ylam
idas
e
+ N
D
ND
N
D
ND
N
D
N
D
-D
-Lac
tosi
dase
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
Le
ucin
e ar
ylam
idas
e N
D
ND
+ +
ND
+
+ +
ND
N
D
ND
+
Le
ucin
e am
inop
eptid
ase
ND
+
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
Lysi
ne d
ecar
boxy
lase
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
-D
-Man
nosi
dase
N
D
ND
N
D
ND
ND
N
D
N
D
ND
N
D
ND
N
D
-M
anno
sida
se
ND
ND
N
D
ND
+k
N
D
O
rnith
ine
deca
rbox
ylas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
Ph
osph
oam
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
Py
rogl
utam
ic a
cid
aryl
amid
ase
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
ND
Pyro
lidon
ylar
ylam
idas
e +
+
ND
N
D
ND
Si
alid
ase
(neu
ram
inid
ase)
d
ND
+
ND
s
ND
N
D
N
D
ND
Ala
-phe
-pro
ary
lam
idas
et N
D
+ N
D
ND
N
D
+ N
D
ND
N
D
ND
N
D
+ N
D
-X
ylos
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
-D
-Xyl
osid
ase
ND
N
D
ND
N
D
N
D
ND
ND
N
D
ND
N
D
ND
Chy
mot
ryps
in
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
Hya
luro
nida
se
+ N
D
d N
D
N
D
d
d N
D
N
D
ND
Ure
ase
+h
()
ND
ND
Val
ine
amin
opep
tidas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
A
ceto
in (V
P)
+ h,
u ()
+
+
+
+ +
+
H
2O2
ND
N
D
ND
N
D
d N
D
ND
ND
+ N
D
ND
IgA
1 pr
otea
se
ND
N
D
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
Extra
cellu
lar p
olys
acch
arid
e N
D
ND
N
D
+ d
ND
N
D
Am
ylas
e bi
ndin
g N
D
ND
N
D
ND
d N
D
N
D
ND
+
ND
N
D
CA
MP
reac
tion
N
D
+ N
D
N
D
h (
+)
ND
N
D
ND
Gro
wth
at:
10C
d
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
45
C
+
ND
N
D
ND
N
D
ND
ND
N
D
ND
G
rowt
h in
pre
senc
e of
:
6.
5% w
/w N
aCl
d
ND
N
D
N
D
ND
N
D
ND
N
D
ND
40%
bile
ND
d
N
D
ND
ND
N
D
ND
N
D
ND
N
D
pH
9.6
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
Hem
olyt
ic R
eact
ion
(,
) ,N
H
NH
,,
,,N
H
N
H,
La
ncef
ield
gro
up a
ntig
en
A
NG
B
D
(G)
NG
, F, C
, G, A
N
G
G
NG
, F, C
, G, A
C
N
G
ND
N
D
NG
Pe
ptid
ogly
can
type
x Ly
sA
la1
3 Ly
sSe
rG
ly
Lys
Ala
13 (
Ser)
N
D
Lys
Ala
13
Lys
Ala
Gly
Ly
sTh
rG
ly
Lys
Ala
13
ND
Ly
sTh
rA
la
ND
N
D
ND
M
ajor
cel
l-wal
l car
bohy
drat
e co
nstit
uent
sz
Rha
, Glc
NA
c R
ha, G
al, (
Glc
NA
c)
Rha
, Gal
, Glc
NA
c , G
luci
tol
ND
R
ha, G
lc, G
al, G
alN
Ac
ND
N
D
Rha
, Glc
, Gal
, Glc
NA
c N
D
Rha
, Glc
, Gal
N
D
ND
N
D
Hos
t M
an
Cat
tle, m
ilk
Man
, ani
mal
s A
nim
als
Man
M
an
Ani
mal
s, m
an
Man
M
an
Ani
mal
s (m
an)aa
M
an
Ani
mal
s A
nim
als
Sour
ce o
f dat
aab
19, 2
6, 2
7, 4
0 16
, 19,
40
19, 2
6 23
, 35,
47
1, 2
7, 3
8, 5
1, 5
2, 5
5 58
13
, 19
27, 5
1, 5
4, 5
5 27
, 51,
54,
55
4, 1
2, 2
7, 3
3 25
, 27
62
38
666 FAMILY VI. STREPTOCOCCACEAE
TAB
LE 1
31.
(con
tin
ued)
Char
acte
ristic
S. downei
S. dysgalactiae subsp.
dysgalactiae
S. dysgalactiae subsp.
equisimilis
S. entericus
S. equi subsp. equi
S. equi subsp.
zooepidemicus
S. equinus
S. ferus
S. gallinaceus
S. gallolyticus subsp.
gallolyticus
S. gallolyticus subsp.
macedonicus
S. gordonii
S. hyointestinalis
S. hyovaginalis
S. infantarius
Cel
l siz
e (m
) N
D
ND
N
D
ND
N
D
ND
N
D
0.5
ND
N
D
ND
N
D
ND
N
D
ND
C
atal
ase
Ac
id p
rodu
ctio
n fro
m:
N-A
cety
l-glu
cosa
min
e N
D
ND
N
D
ND
N
D
ND
+
+ N
D
ND
+
+ +
+ N
D
A
doni
tol
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
A
myg
dalin
N
D
ND
N
D
ND
N
D
ND
+
+ N
D
+
+ d
N
D
A
rabi
nose
ND
A
rbut
in
ND
N
D
ND
N
D
ND
N
D
+ d
ND
N
D
d +
+ N
D
ND
Ara
bito
l N
D
ND
N
D
N
D
ND
ND
C
ello
bios
e
ND
N
D
ND
N
D
ND
+
+ N
D
ND
+
+ d
+ N
D
C
yclo
dext
rin
ND
N
D
ND
+
ND
N
D
N
D
ND
N
D
Dex
tran
ND
N
D
ND
N
D
ND
N
D
N
D
ND
N
D
ND
N
D
ND
N
D
ND
Dul
cito
l N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
Er
ythr
itol
ND
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
Escu
lin
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
+
ND
N
D
ND
Fruc
tose
+
ND
N
D
ND
N
D
ND
+
+ N
D
ND
+
+ +
+ N
D
Fu
cose
N
D
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
G
alac
tose
+
ND
N
D
ND
N
D
ND
+
+ N
D
ND
+
+ +
+ N
D
G
entio
bios
e N
D
ND
N
D
ND
N
D
ND
d
ND
N
D
ND
ND
d
N
D
G
luco
nate
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
N
D
2-
Ket
oglu
cona
te
ND
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
5-K
etog
luco
nate
N
D
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
G
luco
se
+ +
+ N
D
+ +
+ +
+ N
D
+ +
+ +
ND
Gly
cero
l
d d
ND
N
D
ND
d
N
D
G
lyco
gen
d +
+ +
d +
+
d
Inos
itol
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
Inul
in
+
d d
ND
d
d
Lact
ose
+ d
d +
+
d +
+ +
+ +
+ +
+
Lyxo
se
ND
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
Mal
tose
+
+ +
+ +
+ +
+ +
+ +
+ +
+ +
M
anno
se
+ N
D
ND
N
D
ND
N
D
+ +
ND
N
D
+ +
+ +
ND
Man
nito
l +
d
dh (+
) +
+
+
M
elib
iose
ND
N
D
N
D
ND
dh
()
d +
h (
+)
ND
d
ND
dh (+
)
Mel
ezito
se
ND
N
D
ND
ND
Met
hyl-D
-man
nosi
de
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
N
D
ND
Met
hyl-D
-glu
cosi
de
ND
N
D
ND
+
+ +
d
+ +
d d
d
-Met
hylx
ylos
ide
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
ND
Met
hyl-D
-xyl
osid
e N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
ND
N
D
Pu
llula
n
ND
N
D
+
+
ND
+
d
d
Raf
finos
e
d
d +
d +
d d
d
R
ham
nose
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
R
ibos
e N
D
+ +
+
+
d
Sa
licin
+
ND
N
D
ND
+
+ +
+ N
D
ND
d
+ +
+ N
D
So
rbito
l
h (+)
d
+
dh
(+)
h (
d)
+
Sorb
ose
d
ND
N
D
ND
N
D
ND
N
D
ND
Star
ch
+
+ +
+ +
d +
ND
+
d
+
d
Sucr
ose
+ +
+ +
+ +
+ +
+ +
+ +
+ +
+
Taga
tose
N
D
ND
N
D
d
Tr
ehal
ose
+ +
+ +
d
d +
+ +
+
+ +
D-T
uran
ose
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
Xyl
ose
ND
N
D
ND
N
D
N
D
ND
Xyl
itol
N
D
ND
N
D
ND
N
D
ND
N
D
ND
H
ydro
lysi
s of:
A
rgin
ine
+
+
+ +
+
+
Escu
lin
d
dh (+
) +
d d
+ +
+ +
+
+h (
)
d
Gel
atin
N
D
ND
N
D
ND
ND
ND
N
D
ND
ND
N
D
GENUS I. STREPTOCOCCUS 667
(con
tin
ued)
Hip
pura
te
dh (
)
+
Star
ch
N
D
+ +
+
ND
+
d
+n
ND
+
Prod
uctio
n of
:o
A
cid
phos
phat
ase
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
+
ND
ND
Alk
alin
e ph
osph
atas
e
+ +
+
+
d
+ +
+
-L
-Ara
bino
sida
se
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
-D-G
alac
tosi
dase
N
D
d
d +
d d
d d
+
-D
-Gal
acto
sida
se
ND
+ +
+ d
d d
+
Phos
pho-
-gal
acto
sida
se
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
+
ND
N
D
ND
-D-G
luco
sida
se
ND
N
D
ND
N
D
ND
N
D
ND
+
+ N
D
d d
ND
N
D
ND
-D-G
luco
sida
se
ND
N
D
ND
+
+ +
+ +
+
ND
d
d
-L-F
ucos
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
+ N
D
ND
N
D
-D
-Fuc
osid
ase
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
-L-F
ucos
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
N
D
-D
-Glu
curo
nida
se
ND
+
+
+ +
+
-Glu
cosa
min
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
+ N
D
ND
N
D
N-
Ace
tyl-
-D-g
alac
tosa
min
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
d N
D
ND
N
D
N-
Ace
tyl-
-D-g
luco
sam
inid
ase
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
N-A
cety
l--D
-glu
cosa
min
idas
e N
D
ND
N
D
ND
+ N
D
+q
-Mal
tosi
dase
N
D
ND
N
D
ND
N
D
ND
N
D
+ N
D
ND
N
D
+ N
D
ND
N
D
-M
alto
sida
se
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
Gly
cyl-t
rypt
opha
n ar
ylam
idas
e N
D
ND
N
D
+ N
D
ND
ND
+
ND
d N
D
-D
-Lac
tosi
dase
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
+ N
D
ND
N
D
Le
ucin
e ar
ylam
idas
e N
D
+ +
+ +
+ +
+ N
D
+ +
ND
+
+r
+
Leuc
ine
amin
opep
tidas
e N
D
ND
N
D
ND
N
D
ND
+
ND
N
D
ND
N
D
+ N
D
ND
N
D
Ly
sine
dec
arbo
xyla
se
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
-D-M
anno
sida
se
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
-Man
nosi
dase
N
D
N
D
ND
d
ND
d
+ N
D
O
rnith
ine
deca
rbox
ylas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
Ph
osph
oam
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
d N
D
ND
N
D
Py
rogl
utam
ic a
cid
aryl
amid
ase
ND
N
D
ND
ND
N
D
ND
N
D
N
D
ND
N
D
ND
N
D
ND
Pyro
lidon
ylar
ylam
idas
e N
D
d
N
D
Si
alid
ase
(neu
ram
inid
ase)
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
N
D
A
la-p
he-p
ro a
ryla
mid
aset
ND
N
D
ND
+
ND
N
D
+ N
D
+ +
+ +
ND
+
+
-Xyl
osid
ase
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
-D-X
ylos
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
N
D
C
hym
otry
psin
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
+ N
D
ND
N
D
H
yalu
roni
dase
N
D
+ +
ND
+
+ N
D
ND
N
D
ND
N
D
ND
N
D
U
reas
e
ND
N
D
V
alin
e am
inop
eptid
ase
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
+
ND
N
D
ND
Ace
toin
(VP
) +
+
+
+ +
+
+ +
H
2O2
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
+
ND
N
D
ND
IgA
1 pr
otea
se
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
Extra
cellu
lar p
olys
acch
arid
e +
ND
N
D
ND
N
D
ND
+ N
D
+ N
D
+
ND
Am
ylas
e bi
ndin
g N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
+ N
D
ND
N
D
CA
MP
reac
tion
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
G
rowt
h at
:
10
C
ND
ND
+
N
D
ND
N
D
ND
45C
+
ND
N
D
+ +
ND
N
D
ND
N
D
Gro
wth
in p
rese
nce
of:
6.5%
w/w
NaC
l
N
D
d
40
% b
ile
d N
D
ND
N
D
+ N
D
ND
N
D
ND
N
D
N
D
ND
pH 9
.6
+
N
D
ND
N
D
ND
N
D
ND
N
D
ND
H
emol
ytic
Rea
ctio
n N
D
,NH
NH
, N
H
,NH
La
ncef
ield
gro
up a
ntig
en
NG
C
C
, G, L
, A
D
C
C
D
NG
D
, NG
D
, NG
D
, NG
N
G, H
N
G
NG
N
G, D
Pe
ptid
ogly
can
type
x Ly
sTh
rA
la
Lys
Ala
13
Lys
Ala
13
ND
Ly
sA
la1
3 Ly
sA
la2
3 Ly
sTh
rA
la
Lys
Ala
23
ND
N
D
ND
Ly
sA
la1
3 Ly
sA
la (S
er)
ND
N
D
Maj
or c
ell-w
all c
arbo
hydr
ate
cons
titue
ntsz
N
D
Rha
, Gal
NA
c N
D
ND
R
ha, G
alN
Ac
ND
N
D
Rha
, Glc
N
D
ND
N
D
Rha
, Glc
, Gal
, Glc
NA
c, (G
alN
Ac)
N
D
ND
N
D
Hos
t A
nim
als
Ani
mal
s M
an, a
nim
als
Ani
mal
s A
nim
als
Ani
mal
s, m
an
Ani
mal
s A
nim
als
Bird
s A
nim
als,
man
D
airy
M
an
Ani
mal
s A
nim
als
Man
So
urce
of d
ataa
b 5,
57
19, 2
2, 4
8, 5
0 19
, 22,
48,
50
49
9, 1
9, 2
2, 2
7 9,
19,
22,
27
21, 4
1 2,
12
11
7, 2
4, 3
4, 3
6, 4
1, 4
5 7,
24,
34,
36,
41,
45
3, 3
0 14
15
36
, 42
668 FAMILY VI. STREPTOCOCCACEAE
TAB
LE 1
31.
(con
tin
ued)
Char
acte
ristic
S. infantis
S. iniae
S. intermedius
S. lutetiensis
S. macacae
S. minor
S. mitis
S. mutans
S. oligofermentans
S. oralis
S. orisratti
S. ovis
S. parasanguinis
S. parauberis
S. pasteurianus
S. peroris
S. phocae
Cel
l siz
e (m
) 0.
61.
0 ~1
.5
0.5
1.0
ND
N
D
1.
0 N
D
0.5
0.75
0.
7 N
D
ND
N
D
0.8
1.0
ND
N
D
0.6
0.8
ND
C
atal
ase
N
D
Ac
id p
rodu
ctio
n fro
m:
N-A
cety
l-glu
cosa
min
e N
D
+ +
ND
+
+ +
+ N
D
+ N
D
+
+ +
ND
+
A
doni
tol
ND
ND
N
D
ND
ND
N
D
ND
N
D
ND
N
D
ND
Am
ygda
lin
d N
D
+ d
d
+ N
D
d d
ND
ND
Ara
bino
se
Arb
utin
+ +
ND
N
D
d
+
+
ND
d
+ N
D
N
D
A
rabi
tol
ND
ND
N
D
ND
ND
ND
ND
Cel
lobi
ose
ND
+
d N
D
+ +
d +
+ N
D
d d
ND
N
D
ND
Cyc
lode
xtrin
ND
N
D
ND
N
D
N
D
d N
D
ND
N
D
D
extra
n N
D
+ N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
N
D
D
ulci
tol
ND
ND
N
D
ND
ND
N
D
ND
N
D
ND
d
ND
N
D
ND
Eryt
hrito
l N
D
N
D
ND
N
D
N
D
N
D
d N
D
ND
N
D
N
D
ND
N
D
Es
culin
N
D
ND
N
D
ND
N
D
ND
+ N
D
N
D
ND
N
D
ND
N
D
ND
N
D
Fr
ucto
se
ND
+
ND
N
D
+ +
+ N
D
ND
+
+ N
D
+ +
ND
N
D
+
Fuco
se
ND
ND
N
D
ND
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
Gal
acto
se
ND
+f
N
D
ND
+
+ +
+ N
D
+ +
ND
+
+ N
D
ND
G
entio
bios
e N
D
+ N
D
ND
N
D
+ N
D
ND
N
D
ND
N
D
ND
N
D
d N
D
ND
G
luco
nate
N
D
N
D
ND
N
D
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
N
D
2-
Ket
oglu
cona
te
ND
ND
N
D
ND
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
5-K
etog
luco
nate
N
D
N
D
ND
N
D
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
G
luco
se
ND
+
+ N
D
+ +
+ +
+ +
+ +
+ +
ND
N
D
+
Gly
cero
l N
D
ND
N
D
d N
D
ND
N
D
ND
G
lyco
gen
+
+
ND
+ +
d
Inos
itol
ND
ND
N
D
ND
ND
N
D
ND
N
D
ND
ND
N
D
Inul
in
d
N
D
d
+
+ N
D
d
Lact
ose
+
+ +
ND
+
+ +
d +
+ +
+ +
+ +
Lyxo
se
ND
ND
N
D
ND
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
Mal
tose
+
+ +
+ +
+ +
+ +
+ +
+ +
+ +
+ +
M
anno
se
ND
+
ND
N
D
ND
+
+ +
+ +
ND
N
D
ND
+
ND
N
D
+
Man
nito
l
+
+
+i
+
+
dh
(+)
Mel
ibio
se
d d
d
+
d
+
ND
Mel
ezito
se
+
ND
d
Met
hyl-D
-man
nosi
de
ND
+
ND
N
D
ND
ND
ND
N
D
ND
N
D
ND
ND
N
D
ND
Met
hyl-D
-glu
cosi
de
d +
ND
ND
ND
ND
+
ND
-Met
hylx
ylos
ide
ND
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
Met
hyl-D
-xyl
osid
e N
D
N
D
ND
N
D
N
D
N
D
ND
N
D
ND
N
D
N
D
ND
N
D
Pu
llula
n d
ND
+
N
D
ND
d
N
D
+ N
D
N
D
ND
N
D
R
affin
ose
+
+ d
d +
d d
+ +
d d
+
Rha
mno
se
ND
ND
N
D
ND
ND
N
D
ND
N
D
ND
ND
N
D
ND
Rib
ose
+
d
d
ND
ND
d
+
Salic
in
ND
+
d N
D
ND
+
d +
d
+ N
D
d +
ND
N
D
Sorb
itol
+j
d
h (d)
+
+
dh (
) dh
(+)
Sorb
ose
ND
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
Star
ch
ND
+
ND
d
k
d
d +
ND
d
d
ND
d
Su
cros
e +
+ +
+ +
+ +
+ +
+ +
+ +
+ +
+
Ta
gato
se
+
ND
d
d
ND
d
ND
d
ND
d
d
ND
Treh
alos
e
+ +
+
+ d
+ d
d +
+ d
+ +
D
-Tur
anos
e N
D
N
D
ND
N
D
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
X
ylos
e N
D
N
D
ND
N
D
ND
N
D
ND
N
D
ND
X
ylito
l N
D
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
N
D
Hyd
roly
sis o
f:
Arg
inin
e
+h (
) +
+ dh
()
d +
+h (d
)
Es
culin
+ +
+ +
+
+
dl
+ +
d +
+
Gel
atin
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
GENUS I. STREPTOCOCCUS 669
(con
tin
ued)
Hip
pura
te
ND
+
ND
ND
dh
(+)
Star
ch
ND
+
ND
d
+ N
D
d +
ND
+
+ N
D
ND
N
D
N
D
Pr
oduc
tion
of:o
Aci
d ph
osph
atas
e N
D
ND
N
D
ND
N
D
ND
d
d N
D
+ N
D
d N
D
ND
N
D
ND
N
D
A
lkal
ine
phos
phat
ase
+
+
ND
d
d
d +
+ +
d
+
-L-A
rabi
nosi
dase
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
N
D
ND
N
D
ND
N
D
-D
-Gal
acto
sida
se
+
ND
dk
d
+ d
d
d +
d +
N
D
-D
-Gal
acto
sida
se
+
+
ND
d
+
+
+
p N
D
Ph
osph
o--g
alac
tosi
dase
N
D
ND
N
D
ND
N
D
ND
d
ND
N
D
d N
D
ND
N
D
ND
N
D
ND
N
D
-D
-Glu
cosi
dase
N
D
ND
+
ND
N
D
+ +
+ N
D
+ N
D
+ +
ND
N
D
ND
N
D
-D
-Glu
cosi
dase
ND
d
+ N
D
ND
+ N
D
N
D
+ d
ND
+
N
D
-L
-Fuc
osid
ase
ND
N
D
N
D
ND
N
D
ND
ND
d N
D
ND
N
D
ND
-D-F
ucos
idas
e d
ND
+
ND
N
D
ND
d
N
D
N
D
ND
d
ND
N
D
N
D
-L
-Fuc
osid
ase
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
N
D
ND
N
D
ND
-D-G
lucu
roni
dase
ND
N
D
+
-G
luco
sam
inid
ase
ND
N
D
ND
N
D
ND
N
D
d
ND
+
ND
N
D
ND
N
D
ND
N
D
ND
N-A
cety
l--D
-gal
acto
sam
inid
ase
ND
N
D
+ N
D
ND
N
D
ND
+
ND
N
D
ND
N
D
ND
N
D
ND
N-A
cety
l--D
-glu
cosa
min
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
N
D
ND
N
D
ND
N
D
N-
Ace
tyl-
-D-g
luco
sam
inid
ase
d N
D
+
ND
N
D
+
ND
+
ND
+ N
D
ND
-Mal
tosi
dase
N
D
ND
N
D
ND
N
D
ND
d
d N
D
+ N
D
ND
N
D
ND
N
D
ND
N
D
-M
alto
sida
se
ND
N
D
ND
N
D
ND
N
D
d
ND
ND
N
D
ND
N
D
ND
N
D
ND
Gly
cyl-t
rypt
opha
n ar
ylam
idas
e
ND
N
D
N
D
ND
d
N
D
+ N
D
d N
D
ND
N
D
-D
-Lac
tosi
dase
N
D
ND
N
D
ND
N
D
ND
d
+ N
D
N
D
ND
N
D
ND
N
D
ND
N
D
Le
ucin
e ar
ylam
idas
e N
D
+ +
ND
N
D
+ N
D
+ N
D
+ +
+ +
+ +
ND
N
D
Le
ucin
e am
inop
eptid
ase
ND
N
D
ND
+
ND
N
D
+ N
D
ND
+
ND
N
D
ND
+
ND
N
D
ND
Lysi
ne d
ecar
boxy
lase
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
-D
-Man
nosi
dase
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
N
D
ND
N
D
ND
N
D
-M
anno
sida
se
N
D
ND
ND
N
D
ND
ND
ND
N
D
+
ND
Orn
ithin
e de
carb
oxyl
ase
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
Phos
phoa
mid
ase
ND
N
D
ND
N
D
ND
N
D
d +
ND
d
ND
+k
N
D
ND
N
D
ND
N
D
Py
rogl
utam
ic a
cid
aryl
amid
ase
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
N
D
ND
N
D
Py
rolid
onyl
aryl
amid
ase
d +
ND
d N
D
N
D
+
Sial
idas
e (n
eura
min
idas
e)
ND
N
D
+ N
D
ND
N
D
d
ND
+
ND
N
D
N
D
ND
N
D
ND
Ala
-phe
-pro
ary
lam
idas
et +
ND
N
D
+ N
D
ND
N
D
ND
N
D
+ N
D
+ N
D
ND
+
+ N
D
-X
ylos
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
N
D
ND
N
D
ND
N
D
ND
N
D
-D
-Xyl
osid
ase
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
ND
N
D
ND
N
D
ND
Chy
mot
ryps
in
ND
N
D
ND
N
D
ND
N
D
+ +
ND
+
ND
ND
N
D
ND
N
D
ND
Hya
luro
nida
se
ND
N
D
+ N
D
ND
N
D
ND
ND
N
D
N
D
ND
N
D
Ure
ase
N
D
N
D
ND
N
D
ND
Val
ine
amin
opep
tidas
e N
D
ND
N
D
ND
N
D
+ +
+ N
D
+ N
D
ND
N
D
ND
N
D
ND
N
D
A
ceto
in (V
P)
+ +
+ N
D
+
+h
(d)
+
H2O
2
ND
N
D
N
D
N
D
+ d
ND
+
ND
N
D
+ N
D
ND
N
D
ND
IgA
1 pr
otea
se
ND
N
D
ND
N
D
ND
N
D
d
ND
+
ND
N
D
ND
N
D
ND
N
D
ND
Extra
cellu
lar p
olys
acch
arid
e N
D
ND
+
ND
+ N
D
d
ND
ND
ND
N
D
Am
ylas
e bi
ndin
g N
D
ND
ND
N
D
ND
N
D
N
D
ND
d
ND
N
D
ND
N
D
CA
MP
reac
tion
ND
+
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
ND
N
D
G
rowt
h at
:
10
C
ND
+
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
45C
N
D
N
D
ND
ND
N
D
d
ND
+
ND
d
ND
N
D
ND
Gro
wth
in p
rese
nce
of:
6.5%
w/w
NaC
l N
D
d N
D
ND
d N
D
ND
h (
+)
N
D
40%
bile
N
D
N
D
ND
+
ND
N
D
ND
N
D
ND
+
ND
d
ND
N
D
ND
pH
9.6
N
D
ND
N
D
ND
ND
N
D
ND
ND
+
ND
N
D
N
D
ND
N
D
Hem
olyt
ic R
eact
ion
NH
,,
NH
, ()
, N
H
La
ncef
ield
gro
up a
ntig
en
ND
N
G
NG
, F, C
N
G, D
N
G
NG
N
G, O
w, K
N
G
ND
N
G
A
ND
N
G (F
, G, C
, B)
NG
D
N
D
NG
, F, C
Pe
ptid
ogly
can
type
x N
D
Lys
Ala
13
Lys
Ala
13
ND
N
D
ND
Ly
s-di
rect
Ly
sA
la2
3 N
D
Lys-
dire
ct
ND
N
D
ND
N
D
ND
N
D
ND
M
ajor
cel
l-wal
l car
bohy
drat
e co
nstit
uent
sz
ND
R
ha, G
lc, G
al, G
lcN
Ac
Rha
, Glc
N
D
Rha
, Glc
N
D
Gal
, Glu
, Rtl
Rha
, Glc
N
D
Glc
, Gal
,Glc
NA
c, G
alN
Ac,
(Rha
), R
tly
ND
N
D
ND
N
D
ND
N
D
ND
H
ost
Man
Fi
sh, m
an
Man
M
an
Ani
mal
s A
nim
als
Man
M
an
Man
M
an
Ani
mal
s A
nim
als
Man
A
nim
als
Man
M
an
Ani
mal
s (se
als)
So
urce
of d
ataa
b 28
17
, 18,
35
1, 2
7, 4
0, 5
1, 5
2, 5
5 36
, 42
4 46
3,
30
6, 1
2, 2
7, 3
0 44
3,
30,
32
61
10
53
9, 6
0 35
, 42
29
9, 4
3
670 FAMILY VI. STREPTOCOCCACEAE
TAB
LE 1
31.
(con
tin
ued)
Char
acte
ristic
S. pluranimalium
S. pneumoniae
S. porcinus
S. ratti
S. salivarius
S. sanguinis
S. sinensis
S. sobrinus
S. suis
S. thermophilus
S. thoraltensis
S. uberis
S. urinalis
S. vestibularis
Cel
l siz
e (m
) N
D
0.5
1.25
N
D
0.5
0.8
1.0
ND
0.
820
.98
0.5
GENUS I. STREPTOCOCCUS 671
(con
tin
ued)
Hip
pura
te
+h (
)
h (
d)
d
+
Star
ch
ND
+ d
ND
N
D
+ d
ND
N
D
+
Prod
uctio
n of
:o
Aci
d ph
osph
atas
e N
D
ND
N
D
ND
+
d N
D
ND
N
D
ND
+
ND
Alk
alin
e ph
osph
atas
e d
+
ND
+
dk
d
+
-L
-Ara
bino
sida
se
ND
N
D
ND
N
D
ND
ND
N
D
ND
N
D
ND
N
D
-D
-Gal
acto
sida
se
d +
N
D
d
+
d
-D
-Gal
acto
sida
se
d +
N
D
d d
d +
p
+
Phos
pho-
-gal
acto
sida
se
ND
N
D
ND
N
D
d
ND
N
D
ND
N
D
ND
N
D
ND
N
D
-D
-Glu
cosi
dase
+
+ N
D
ND
+
N
D
+ N
D
N
D
ND
+
d
-D-G
luco
sida
se
dd
d N
D
ND
+
d +
N
D
+
ND
+
-L-F
ucos
idas
e N
D
d N
D
ND
N
D
N
D
N
D
ND
-D-F
ucos
idas
e N
D
d N
D
ND
d
d N
D
N
D
ND
N
D
ND
N
D
-L-F
ucos
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
-D-G
lucu
roni
dase
dd
d N
D
+
+ +
-G
luco
sam
inid
ase
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
N-
Ace
tyl-
-D-g
alac
tosa
min
idas
e N
D
+ N
D
ND
N
D
N
D
N
D
ND
N
D
N-A
cety
l--D
-glu
cosa
min
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N-A
cety
l--D
-glu
cosa
min
idas
e d
d N
D
ND
d
+
N
D
-Mal
tosi
dase
ND
N
D
ND
+
+ N
D
ND
N
D
ND
N
D
ND
N
D
ND
-Mal
tosi
dase
N
D
ND
N
D
ND
d
N
D
ND
N
D
ND
N
D
ND
N
D
ND
Gly
cyl-t
rypt
opha
n ar
ylam
idas
e N
D
+
ND
d N
D
d
ND
d
-D-L
acto
sida
se
ND
N
D
ND
N
D
d d
ND
N
D
ND
N
D
ND
N
D
ND
N
D
Le
ucin
e ar
ylam
idas
e +
d +
ND
+
ND
+
ND
+
+ +
+ +
ND
Leuc
ine
amin
opep
tidas
e +
ND
N
D
ND
+
+ N
D
ND
N
D
ND
+
+ +
ND
Lysi
ne d
ecar
boxy
lase
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
-D-M
anno
sida
se
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
N
D
-M
anno
sida
se
N
D
N
D
d N
D
O
rnith
ine
deca
rbox
ylas
e N
D
ND
N
D
ND
N
D
ND
+
ND
N
D
ND
N
D
ND
Phos
phoa
mid
ase
ND
N
D
ND
N
D
+ d
ND
N
D
ND
N
D
ND
N
D
ND
N
D
Py
rogl
utam
ic a
cid
aryl
amid
ase
d N
D
N
D
N
D
ND
N
D
d N
D
ND
d
+
Py
rolid
onyl
aryl
amid
ase
dh (
) dh
()
h (
+)
ND
+ +
Sial
idas
e (n
eura
min
idas
e)
ND
+
ND
N
D
ND
ND
N
D
ND
N
D
ND
A
la-p
he-p
ro a
ryla
mid
aset
+ N
D
+ N
D
+ +
N
D
+ N
D
+ +
+
-X
ylos
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
-D-X
ylos
idas
e N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
ND
Chy
mot
ryps
in
ND
N
D
ND
N
D
+ d
ND
N
D
ND
N
D
ND
N
D
N
D
H
yalu
roni
dase
N
D
+ N
D
ND
N
D
d
ND
N
D
ND
N
D
Ure
ase
ND
d
N
D
+
V
alin
e am
inop
eptid
ase
d N
D
ND
N
D
+ d
ND
N
D
ND
ND
N
D
N
D
A
ceto
in (V
P)
dh (+
) +
dh (+
)
+ +
+
+ +h
(d)
+ +h
(d)
H
2O2
ND
+
ND
+
ND
+
ND
N
D
ND
N
D
ND
+
Ig
A1
prot
ease
N
D
+ N
D
ND
+ N
D
ND
N
D
ND
N
D
ND
N
D
ND
Extra
cellu
lar p
olys
acch
arid
e N
D
N
D
+ +
+ N
D
+
ND
N
D
ND
A
myl
ase
bind
ing
ND
N
D
ND
N
D
d
ND
ND
N
D
ND
N
D
ND
CA
MP
reac
tion
ND
N
D
+ N
D
ND
N
D
ND
N
D
ND
N
D
ND
N
D
Gro
wth
at:
10
C
ND
ND
ND
45
C
ND
d
d d
ND
d
+
ND
Gro
wth
in p
rese
nce
of:
6.
5% w
/w N
aCl
dd,h (+
)
ND
d
+
h (+)
+
40%
bile
N
D
N
D
d d
d +
d d
N
D
d N
D
pH 9
.6
ND
ND
ND
N
D
N
D
N
D
ND
H
emol
ytic
Rea
ctio
n
NH
N
H,,
()
NH
, (
)v
,NH
N
H
La
ncef
ield
gro
up a
ntig
en
ND
N
G
E, P
, U, V
N
G
NG
, K
NG
, H
NG
N
G
R, S
, T, N
G
NG
N
G
NG
N
G
NG
Pe
ptid
ogly
can
type
x N
D
Lys
Ala
2 (Se
r)
Lys
Ala
24
Lys
Ala
23
Lys
Ala
24,
Lys
Thr
Gly
Ly
sA
la1
3 N
D
Lys
Thr
Ala
Ly
s-di
rect
(Lys
Ala
12)
Lys
Ala
23
ND
Ly
sA
la1
3 N
D
ND
M
ajor
cel
l-wal
l car
bohy
drat
e co
nstit
uent
sz
ND
G
lc, (
Gal
), G
alN
Ac,
Rtl
ND
R
ha, G
al, G
ro
Rha
, Glc
, Gal
, Gal
NA
c R
ha, G
lc, G
lcN
Ac
ND
R
ha, G
lc, G
al
Rha
, Glc
, (G
al),
(Gal
NA
c)
ND
N
D
Rha
, Glc
, Glc
NA
c N
D
ND
H
ost
Ani
mal
s M
an
Ani
mal
s M
an, a
nim
als
Man
M
an
Man
M
an, a
nim
als
Ani
mal
s, m
an
(Dai
ry p
rodu
cts)
A
nim
als
Ani
mal
s M
an
Man
So
urce
of d
ataa
b 16
19
, 27,
40
8, 9
12
, 33
3, 1
9, 3
0, 4
0 3,
19,
26,
27,
30,
40
60
6, 1
2, 3
3 19
, 31
20, 3
9, 5
5 15
8,
9, 1
9, 2
6, 5
9 9
56
672 FAMILY VI. STREPTOCOCCACEAE
a Sym
bols
: +, >
85%
pos
itive
; d, d
iffer
ent s
train
s giv
e di
ffer
ent r
eact
ions
(16
84%
pos
itive
); ,
015
% p
ositi
ve; w
, wea
k re
actio
n; N
D, n
ot d
eter
min
ed.
b Aci
d pr
oduc
tion
from
car
bohy
drat
es o
ften
wea
k an
d di
ffic
ult t
o in
terp
ret.
c Isol
ates
bec
ome
cata
lase
-neg
ativ
e on
subc
ultu
ring.
d S
train
s fro
m b
ovin
e ge
nita
l tra
ct d
iffer
from
thos
e fr
om o
ther
sour
ces.
e Rea
ctio
n va
ries a
ccor
ding
to te
st k
it us
ed.
f Rep
orts
var
y.
g Som
e st
rain
s giv
e w
eak
reac
tions
. h C
hara
cter
istic
scor
e di
ffer
s fro
m F
ackl
am (2
002)
, with
the
latte
rs s
core
giv
en in
par
enth
eses
. i A
cid
prod
uctio
n de
laye
d.
j Aci
d pr
oduc
tion
slow
. k W
eak
reac
tion.
l S
ome
repo
rts n
egat
ive.
mSl
ow re
actio
n.
n Pos
itive
or w
eak
reac
tion
on M
lle
rH
into
n ag
ar.
o Pre
form
ed e
nzym
e ac
tiviti
es d
etec
ted
are
depe
nden
t on
the
test
form
at (s
ubst
rate
) use
d an
d m
ay b
e af
fect
ed b
y cu
lture
con
ditio
ns. I
ndiv
idua
l enz
yme
activ
ities
repo
rted
in th
e lit
erat
ure
may
ther
efor
e va
ry.
p Sub
stra
te-d
epen
dent
. q R
esul
ts m
ay v
ary.
r R
eact
ions
freq
uent
ly w
eak.
s C
onfli
ctin
g re
sults
repo
rted.
t A
la-p
he-p
ro, a
lany
l-phe
nyla
lany
l-pro
line
[ary
lam
idas
e].
u Rep
orte
d re
sults
diff
er.
v -H
emol
ysis
may
be
obta
ined
on
hors
e bl
ood-
cont
aini
ng a
gar.
wO
ant
igen
pre
sent
in b
iova
r 1 st
rain
s. x A
bbre
viat
ion
of p
eptid
ogly
can
type
acc
ordi
ng to
Sch
leife
r and
Kan
dler
(197
2).
y Rib
itol t
eich
oic
acid
and
pho
spho
chol
ine
pres
ent.
z Gal
(gal
acto
se);
Glu
(glu
cose
); G
alN
Ac
(N-a
cety
lgal
acto
sam
ine)
; Glc
NA
c (N
-ace
tylg
luco
sam
ine;
Gro
(gly
cero
l); R
ha (r
ham
nose
); R
tl (r
ibito
l).
aaO
ccas
iona
lly m
an.
abD
ata
obta
ined
from
: 1. A
hmet
et a
l. (1
995)
; 2. B
aele
et a
l. (2
003)
; 3. B
eigh
ton
et a
l. (1
991)
; 4. B
eigh
ton
et a
l. (1
984)
; 5. B
eigh
ton
et a
l. (1
981)
; 6. B
eigh
ton
et a
l. (1
991)
; 7. B
rook
er e
t al.
(199
4); 8
. Col
lins e
t al.
(198
4a);
9. C
ollin
s et a
l. (2
000)
; 10
. Col
lins e
t al.
(200
1); 1
1. C
ollin
s et a
l. (2
002)
; 12.
Coy
kend
all (
1977
); 13
. Dev
riese
e a
l. (1
986)
; 14.
Dev
riese
et a
l. (1
988)
; 15.
Dev
riese
et a
l. (1
997)
; 16.
Dev
riese
et a
l. (1
999)
; 17.
Eld
ar e
t al.
(199
4); 1
8. E
ldar
et a
l. (1
995)
; 19.
Fac
klam
(2
002)
; 20.
Far
row
and
Col
lins (
1984
b, 1
984a
); 21
. Far
row
et a
l. (1
984)
; 22.
Flin
t et a
l. (1
999)
; 23.
Han
dley
et a
l. (1
991)
; 24.
Har
die
(198
6a, 1
986b
); 25
. Har
die
and
Whi
ley
(199
5); 2
6. K
awam
ura
et a
l. (1
998)
; 27.
Kaw
amur
a et
al.
(199
8); 2
8.
Kili
an e
t al.
(198
9b);
29. K
ilppe
r-B
lz
and
Schl
eife
r (19
87);
30. K
ilppe
r-B
lz
et a
l. (1
985)
; 31.
Kra
l and
Dan
eo-M
oore
(198
1); 3
2. O
saw
a et
al.
(199
5); 3
3. P
ier a
nd M
adin
(197
6); 3
4. P
oyar
t et a
l. (2
002)
; 35.
Rob
inso
n et
al.
(198
8); 3
6.
Rur
angi
rwa
et a
l. (2
000)
; 37.
Sch
leife
r et a
l. (1
991)
; 38.
Sch
leife
r and
Kilp
per-
Bl
z (1
987)
; 39.
Sch
lege
l et a
l. (2
003)
; 40.
Sch
lege
l et a
l. (2
000)
; 41.
Ska
ar e
t al.
(199
4); 4
2. T
ong
et a
l. (2
003)
; 43.
Tsa
kalid
ou e
t al.
(199
8); 4
4. V
anca
nney
et a
l. (2
004)
; 45.
Van
dam
me
et a
l. (1
999)
; 46.
Van
dam
me
et a
l. (1
996)
; 47.
Vel
a et
al.
(200
2); 4
8. V
ieira
et a
l. (1
998)
; 49.
Whi
ley
and
Bei
ghto
n (1
991)
; 50.
Whi
ley
and
Bei
ghto
n (1
998)
; 51.
Whi
ley
et a
l. (1
990a
, 199
0b);
52. W
hile
y et
al.
(199
9); 5
3.
Whi
ley
and
Har
die
(198
8); 5
4. W
hile
y et
al.
(198
8); 5
5. W
illco
x et
al.
(200
1); 5
6. W
illia
ms a
nd C
ollin
s (19
90a)
; 57.
Woo
et a
l. (2
002)
; 58.
Zhu
et a
l. (2
000)
; 59.
Col
lins e
t al.
(200
4).
TAB
LE 1
31.
(con
tin
ued)
GENUS I. STREPTOCOCCUS 673
TAB
LE 1
32.
Bio
chem
ical
test
s di
ffer
enti
atin
g -h
emol
ytic
str
epto
cocc
al s
peci
es (
Pyog
enic
gro
up p
lus
An
gin
osus
gro
up)a
Ch
arac
teri
stic
S. pyogenes
S. agalactiae
S. canis
S. didelphis
S. dysgalactiae
subsp. dysgalactiae
S. dysgalactiae
subsp. equisimilis
S. equi subsp. equi
S. equi subsp.
zooepidemicus
S. iniae
S. phocae
S. porcinus
S. anginosus
S. constellatus
subsp. constellatus
S. constellatus
subsp. pharyngis
S. intermedius
Cat
alas
e
+b
Aci
d pr
oduc
tion
from
:
Gly
coge
nd
c
d+
++
d
ND
Man
nit
ol
d
+
d
dd (
)
R
affi
nos
e
d
d
R
ibos
e
++
++
+
++
++
ND
Sorb
itol
d
+
+
Sucr
ose
++
+N
D+
++
++
d
+N
DN
D+
Tr
ehal
ose
++
d+
++
d
+
++
d+
+H
ydro
lysi
s of
:
Arg
inin
e+
++
dd (
+)+
++
++d
()
+
++
++
E
scul
ind
+
d
dd
d+
dd
(+)
++
++
H
ippu
rate
+
+
dd
()
d,f
St
arch
++
+
ND
Prod
uctio
n of
:e
-D-g
alac
tosi
dase
d
dd
N
D
-D
-gal
acto
sida
se
-+
+
ND
d
+
+
-D-g
lucu
ron
idas
ed
dd
++
++
+N
D
d
Py
rolid
onyl
aryl
amid
ase
+
+
d (
+)
A
ceto
in (
VP
)
+d,f
()
dd (
+)+
++
+C
AM
P re
acti
on
+d
(+)
+
+
Lan
cefi
eld
grou
p an
tige
nA
BG
NG
CC
, G, L
, AC
CN
GN
G, F
, CE
, P, U
, VN
G, F
, C, G
, AN
G, F
, C, G
, AC
NG
, F, C
Hos
tM
anM
an,
anim
als
An
imal
s,
man
An
imal
sA
nim
als
Man
, an
imal
sA
nim
als
An
imal
s,
Man
Fish
, an
imal
s,A
nim
als
An
imal
sM
anM
anM
anM
an
a Sym
bols
: +, >
85%
pos
itiv
e; d
, dif
fere
nt s
trai
ns
give
dif
fere
nt r
eact
ion
s (1
684
% p
osit
ive)
; , 0
15%
pos
itiv
e; w
, wea
k re
acti
on; N
D, n
ot d
eter
min
ed.
b Iso
late
s be
com
e ca
tala
se-n
egat
ive
on s
ubcu
ltur
ing.
man
c Som
e st
rain
s gi
ve w
eak
reac
tion
s.d C
har
acte
r sc
ore
diff
ers
from
Fac
klam
(20
02)
wit
h th
e la
tter
s s
core
als
o fo
llow
ing
in p
aren
thes
es. A
n a
ddit
ion
al, u
sefu
l tes
t acc
ordi
ng
to F
ackl
am (
2002
) is
sen
siti
vity
to b
acit
raci
n fo
r S.
pyo
gene
s an
d S.
pho
cae
only
e Pre
form
ed e
nzy
me
acti
viti
es d
etec
ted
are
depe
nde
nt o
n th
e te
st fo
rmat
(su
bstr
ate)
use
d an
d m
ay b
e af
fect
ed b
y cu
ltur
e co
ndi
tion
s. I
ndi
vidu
al e
nzy
me
acti
viti
es r
epor
ted
in th
e lit
erat
ure
may
ther
efor
e va
ry.
f Rep
orte
d re
sult
s di
ffer
.
674 FAMILY VI. STREPTOCOCCACEAE
TABLE 133. Characteristics of Streptococcus species: oral streptococcal species groupsa,b
Anginosus group Mutans group
Characteristic S. a
ngin
osus
S. c
onst
ella
tus
subs
p. c
onst
ella
tus
S. c
onst
ella
tus
subs
p. p
hary
ngis
S. in
term
ediu
s
S. c
rice
ti
S. d
evri
esei
S. d
owne
i
S. fe
rus
S. m
acac
ae
S. m
utan
s
S. o
risr
atti
S. r
atti
S. s
obri
nus
Acid production from: Amygdalin + d + d + + ND + + d + + Arbutin + d + + + + ND d ND + + + Glycogen ND + + Inulin + d + d + + d d Lactose + d + + d + + + ND + + d d Mannitol dc () + + + dc (+) + + + + Melibiose d + + d d + d Raffinose d + + d + + + + Ribose ND ND ND Sorbitol + + c (+) dc (+) +b + + dc (+) Trehalose + d + + d + + + + + + + +Hydrolysis of: Arginine + + + + + Esculin + + + + dc (+) ND + + + + + dc (+) Hippurate + ND ND Production of:e
Acetoin (VP) + + + + + + + + + + + + N-Acetyl--d- glucosaminidase
d + ND ND ND ND ND ND
Alkaline phosphatase
+ + + + ND - d ND ND
Extracellular polysaccharide
+ ND + + + + + +
-d-Galactosidase d + + ND ND ND ND -d-Glucosidase d + + + ND ND ND + ND + ND ND + -d-Glucosidase + + d ND + ND + ND + ND ND -d-Fucosidase + + ND ND ND ND ND ND ND Urease Lancefield group antigen
NG, F,C, G, A
NG, F,C, G, A
C NG, F, C NG ND NG NG NG NG A NG NG
Host Man Man Man Man Animals (man)j
Animals Animals Animals Animals Man Animals Man, animals
Man, animals
aSymbols: +, >85% positive; d, different strains give different reactions (1684% positive); , 015% positive; w, weak reaction; ND, not determined.bAcid production is slow.cCharacter score differs from Facklam (2002), with the latters score given in superscript.dSome reports negative.ePreformed enzyme activities detected are dependent on the test format (substrate) used and may be affected by culture conditions. Individual enzyme activities reported in the literature may therefore vary.fSubstrate-dependent.gSubstrate-dependent.hSubstrate-dependent.iO antigen present in biovar 1 strains.
BERGEYS MANUAL OFSystematicBacteriology IIISecond EditionISBN: 9780387950419PrefaceContents