SANTIAGO ORTIZ and JUAN RODRIGUEZ-OUBI17~A

Departamento de Bioio~ Vexetal, Laboratorio de ~ Facultade de Farmacia, Universidade de Santiago de Compostela, 15706 Santiago de Compostela, Galicia, Espafia

Synopsis of the Rupicolous Vegetation of Galicia (North-western Iberian Peninsula)

KEYWORDS

Ph~csodolo~, Phytogeography, Climate, Asp/en/aea v~oman/J, Th/asp~a roo~m~fo~, ~ ~vo~, New ~ Peninsula, Typification

ABSTRACT

In this paper we present a synopsis of the rupicolous vegetation found in Galicia, according to the Zflrich- MontpeRier School. The climate of this territory, an area of 29,439 km 2 situated in the Northwest of the Iberian Peainsul& is largely Atlantic European beca3min 8 subhumid Mediterranean with a Central European tendency in the east of the region. There is an appreciable summer drought but annual rainfall is high: ombroclime is thus hyperhumid or humid br subhumid in the Southeast. Biogeographically most of the area is Euros~rian eareept part of the Southeast, which is Mediterranean. Acidic rocks (granites, 8neissic, schists and slates) ~ i n : t t ~ ; there are also smaller areas of limestone, gabbros, and metabasie and ultrabasic rocks, some of this

type are serpentine. In this synthesis, we report 17 associations and 16 subassociations as well as 3 communities of unknown

swntaxonomy. Of these, 1 association and 9 subassociations are de.son'bed as new syntaxa and the names of 3 associations are corrected. The principal ecological conditions that influence the composition and distn'bution of these associations and its biogeographic and bioclimafic characteristics are presented in two ~,mm~riTinZ tables. Florisfic differences between the associations included in this synopsis are summarized in a synoptic table.

INTRODUCTION

The rupicolous vegetation of Galicia (tan be divided into two groups. The first, non-halophilic, group can be shown to be closely related to the rupicolous vegetation of the siliceous substrata of the Mediterranean Carpetano-Iberian-Leonesean Province and,

Podia Gea3bot. Phytotax., Praha, 28:15-49

16 FOHA GBOBOTANICA ET rH~TOTAXONO~CA, 28, 1993

to a lesser extent, to the caldphile vegetation of the Cantabrian Mrs. The second, halochasmophytic group is closely related m cliff vegetatiorl found on the southern Atlantic coast of Europe, for example in Britain, Ireland and Cornwall (FERN~'q'DEZ PRIETO and LOIDI 1984) and especially to that found on the Cantabrian coast. However, the vegetation of the western cllf~ of Galicia is already more closely related to that of the Portuguese coast.

The rupicolous vegetation has been partially studied in various phytosociological works, among which those of BELLOT (1951a, b, 1966), C.ASASEC.A (1959), RIVAS-MART~EZ (1969), FERNANDEZ PRmTO (1983), F'F.RNAN~EZ PRIMO and I.OIDI (1984), J. G u m ~ (1984), Og'nz (1986), F E ~ E Z PRmTO et al. (1986), IZCO and Og'nz (1987), F E g N A N D E Z - ~ et al. (1987), P. G u m ~ (1989) and I~REz-C.ARRO et al. (1989) should be mentioned.

In this paper we present a synopsis of the mpicolous vegetation of Galicia, whichis the result of field Work carried out by the authors over the last few years and based on a synthesis of data from the previous studies. The recognized communities are classified in a syntaxonomic synopsis.

MATERIAL AND METHODS

STUDY AgEA

The study region covers an area of 29,439 kin2 and is situated in the northwest of the Iberian Peninsula, in Spain (Fig. 1).

F'~ I. Location of Galicia in the Iberian Peninsula

Following the classification of ALLLr~ (1966), climate is largely European Atlantic becoming subbnmid Mediterranean with a Central European tendency in the east of the region ( ~ , , ~mA et al. 1983). Following the classification of RIV~S-MARTI~Z (1987), six principal bioclimatic belts are distinguished in the region: Colline (with a very characteristic Thermocolline Subbelt), Montane and Subalpine belts in the area pertaining to the Eurosiberian region, and Meso-, Supra-, and Oromediterranean belts in the area corresponding to the Mediterranean region (Fig. 2). These belts are defined by annual average temperature (T), average minimum temperature of the coldest month (m), average

ORTIZ AND RODRIGUEZ-OUBII~ SYNOPSIS OFTHE RUPICOLOUS VEGETATION 17

F'~ 1 Bioclimalic map of Galicia (Bioclimatic . . i t~ after RIVAZ-MARItNEZ 1987)

J Enrosibedan Reg~.

Colline Belt

Montane Belt

m

I-7-1

Thermocolline Subbelt

Eucolline Subbelt

Uppercolline Subbelt

Midmo-J-ne Subbelt

Uppermoatane Subbek

Subalplne Belt

Mediterranean Region

E

Lower Subalpine Subbelt

Mesomediterranean Belt

Supramediterranean Belt

Oromediterrane, an Belt

18 FOLXA OEOE~'rANICA ET PHYrO'rAXONOMICA, 28, 1993

maximum temperature of the coldest month (M), and heat index [ltffi (T+m+M)xl0] (RIVAS-MAR~Z, op. cit.). Annual rainfall, which is high in most of the region despite an appredable summer drought, determines an ombroclimate which varies from the hyperhumid to humid becoming subhumid in the Southeast.

Both biogeographical regions of continental Europe occur in Galicia: the Eurosiberian region (most of Galicia) and the Mediterranean region (a considerable area in the Southeast). Both regions have been subdivided by R_rVAS-MARTI~Z (1987) into Provinces, sectors, and subsectors. Those referring to Galicia are shown in Fig. 3.

The relief of the studied region is very uneven with a continuous succession of hills and valleys. The highest altitude is that of lena Trevinca Sur (2,124 m) but most of the areas are of low (around 400 m) or medium altitude (800-1,000 m) except on the eastern border of the region. The predominant substrata are of an acidic type, especially granites, gneissic schists and clay schists with a acid or neutral pH. Limestone, gabbro, metabasic and ultrabasic siliceous rocks also occur over smaller areas. This last type includes serpentine rocks which have a high level of heavy metals and harbour a very peculiar type of vegetation.

PHYIDSOCIOI.OGICAL INVESTIGATIONS

For the study of the rupicolous vegetation of Galicia, we have followed the method of the Zfirich-Montpelller School (BRAUN-BLANOUET 1964). The syntaxonomic nomenclature followed is, for the most part, that proposed by DIAz-Go~.~J.~z (1989). For the description of new syntaxa, corrections, etc. the "Code of Pbytosociological Nomenclature" ( B ~ et el. 1986) has been followed.

The nomenclature of the taxa present in the relev6s is in accordance with that of Flora Europaea (TurIN et el. 1964-1980) or (in the case of taxa included in the voluraes published to date) the Flora Iberica (CAsTROV~O et al. 1986-1990), with the: exception of the following: Adenocarpus complicatus (L) GAY subsp./a/nz//CAS'~OVIE~O, Agrost/s t//en/NIETO F~Ia~R et ~ s m o v ~ o , Agrost/s truncatu/a PAre. subsp, dur/ad (Bo~s. et REtrma ex WllJ.x.) ASa~RSON et GRABBI~_~, Anarrh/num x /ntermed/um SIMON, Ant/rrh/num meonanthum H o ~ s . et ~ subsp, salcedoi ~ z , Arme~ pubigera (DESF.) Bolss. subsp.pubigera, Armeriapubigera (D~sF.) Bolss. subsp, depilata (B~mcm) F. ~ et LomI, Coincya pseudoerucastrum (BROT.) GREUTER et BUP.DET subsp, setigera (I.ANGE) GREtrtr~ et Btn~ET, ~ oromeditetraneus RrVAS-MART~.Z, DIAz-GoNzAL~Z, F. thtmTO, LoroI et

Dryopteris oreades FOMIN, Ga//um ~ WEIGEL subsp. ~ KLIPHUIS, Hieracium gallicianum PAU, H/erac/um pa///dum Brc.-BERN. subsp, gran/t/aon (SCH.-Bw.) ZAHN, Leontodon farinosus MERINO et PAu, Leucanthemopsis paUida (MnL~R) HEYWOOD subsp, flaveo/a (HoFFMANNS. et I_ZNK) LADERO et VELASCO, Leucanthemum ga//ae/cum Ron~Gt~z-Otam% et Om~., L/nar/a saxat///s (L) ~ subsp, g/abrescens (L~GE) L/thodora dlffusa (LAG.) J.M. JOHNSTON, Lithodora prostrata (LoIsEL.) GRISEB., Plantago a/p/ha L vat. er/opoda WILLr~ et L~GE, Poa/eg/onens/~ (LAINz) FERNAt,mEZ CASAS et LA~Z, Rhamm~ pwn//a Ttya~ subsp. /eg/onends (Ro'mM.) LAI~z, Rumex acetosa L subsp. bifformis (I.~GE) VALD~S B~SX~_JO et CAS~OV~O, Santolina rosmarinifolia L subsp. semidentata ( H o ~ . et LINK) VALD~S BERMEJO, Sesamoide~ minor (I.ANGE) O. KUIr Sesamoides suffmticosa (I_.~GE) O. KtmTZK S//ene un/flora Ro~t, Thlaspi alpestre L subsp. mednoi I.AINZ.

OIU' IZ AND R O D i t I G ~ SYNOIF~,S OF TEIE RUIRICOLOUS VEOETATION 19

F~. 3. ~ mpot GaV~a

. I tO ~ U% I

. . . . . , - ' - " -7 " ' , - -

~, <~ ~" . . . . ~ j ' ~ , j . ~ - ' ~

(~ .,.--~-'~-~. _ ~ . / ," L ,."~'- _ : ~ P / , ~ ' ~ . . ~

,

- - - Sr162 bomKl~ . . . . . . . Su~__~ boundm7

]L~ ix~ Sector SulxMx~

-~acsc

G z l 6 m - ~ . a n

G z l 6 c ~ P ~ p ~

L ~ t s n o - ~

Northern Galako-Ama.ian

C.,o=poa=~ l .uccm/aa

Mieeaah= j ~ o 0

Naviaao-Aacaremiaa

Oremaa

MarlSato-Saoalxhm~

Tralg~o,,t~mn

2a 21)

2c 2d

3

4a

41)

5

2 0 FOHA OEODO'rAN]CA Err rh'YrOTAXONOM~CA, 28, 1993

Table 1. Principal ecological charmeri~a~ that influence the composition and distn~utlon of the studied comml,n;tle~. The data in this table are bas~ m the authors own ~ and those ~ ~ a ~ ~ m the te~

Ecoloocal c h a r m e r t ~

Community Sea So~ Rock Root type spray nitrogen mobility

O C h m o - ~ .z~,i~rae Spogdav/o-Anno/aUm d e ~ ! ~ C.a/endu~Pa~/aum ~_a~'_~ Par~taT~um ~ u C ~ , a , a : a ~ r ~ caou~ ? ~ c.~:_~,n community ~ D e m U / e a , m ~ LOtat/o-Osei/ant~utm ~hae/

Phasnak,-Rum/caum/ndun~

no hiSh n o h loh

no hloh no low no low no low

no low no low

no low

P ~ ~mdO~onx no low Sa~lraSe~n ~ no ~ aovst /cae no low Ou~mtIw-Asp/tn/aum conmnense no low Oy~oyammo-Z~vto~um abb, e ~ m Ro CgNwyammo~//oumm go3~nae no low Ruma =Out~oa~ c ~ , - ~ i t y no low Sesamo/do-S//omUm ~ no low

cOof~l'whtto~ f ~ t l s

c~solidated siliceoes mnsolidated siliceous mmolldated ~i~ous cemolidated si]keous mesolidated siliceous conmlid~ed siticeoes mnsolidated siliceous mesolidated siIkeous comolidated ~lkeous consolidated si]iceoes o r n o t

~ ~ siliceous or not comolidated calcareous mesolidated calcareous consolidated calcareous coasolidated eltrabasic mobile silkeous mobile si]iceous mobile sificeous mobile ,u'li~ous mobile metabasic

RESULTS

ECOLOGY AND BIOGEOGRAPHY OF THE GALICIAIq RUPICOLOUS COMMUNITIES

The principal ecological conditions which influence the florisfic composition and distribution of the studied communities and their biogeographic and bioclimatic characteristics are presented in Tabs. 1 and 2. The first two associations in the Tab. 1 are identical with respect to the four ecological characteristics considered, but are clearly distinct in biogeographical affinities (Tab. 2). Ca/endu~Par/etat/emm juda/cae, occurring in the nitrogen-rich soils typical of seabird nesting colonies, is clearly differentiated from the other syntax& Patietatietum judaicae, ~mbalario-Trachelietum caendei and the Pob~onum cap/tatum community again share the four ecological characteristics, and all occur in the

OI~'IZ AND RODP-JGUEZ=O~ SYNOPSIS OF THE RUPICOLOUS VEGETATION 21

Thermocolline Subbelt of the Mineasian subsector. The differences between them axe solely floristic. In isolated areas in the North of the subsector, L3ymba/ar/o-Trache//etum caende/is constituted by the largely Mediterranean and calcicolous taxon Trache//um caendeum L ; in the extreme Southwest of Galicia, Potygonum cap/tatum D. DoN, a neophyte of Himalayan origin, enters assemblage of Parietarietum j:~__~__'_cae species constituting the Polygonum cap/tatum comm~mity.

The next four associations are not differentiated by the four ecological characteristics given in Tab. 1, but clearly have different geographical distributions. The same applies m P/ug, na/o-Rum/cetwn /ndurat/ and Se.samo/do-Ananh/nenmz dur/m/m', which both occur on embankments and rocky slopes or on consolidated bare rock, almost always in wide soil-fiUed crevices.

The next three associations in the table are identical in terms of the characters shown in the two tables, but they differ in other ecological features. Whilst all of them are calcicolous, Pe~rocoptidetwn grandiflorae generally colonizes rock ledges, Saz/fragetum tr/furcatae narrow crevices, and Asplenio-Cheilanthetum acrosticae wide soft-filled crevices in the warmer areas of Galicia.

Cheilantho-Asplenietum corunnense is the only community to occur on ulU'abasic consolidated rocks. The next four associations occur on screes in siliceous areas; they display some biogeographical differences, but they differ mainly in their preference for rock size. Cryptogrammo-Dryoptetideaan abbreviatae occurs in sites characterized by large quartz blocks, Sesamoido-Silenetum gayanae on screes comprised of small fragments of slate, and the Rumez suffnaicosus community and Cryptogtwnmo-Silenetwn gayanae in areas of medium-size blocks of either quartz or slate. The latter community generally occurs in soils with higher moisture levels than those colonized by the community of Rumex su09ut/cosus.

Finally, the Rum~ scutatus subsp, gallaecicus community, is clearly differentiated from the other syntaxa by being limited to metabasic sites.

Table l Biogeographical and bioellm~ric characteristics of the studied communities. The data in this table are

based on the authors own observations and those of the authors cited in the text.

Community S~ctor

Biogeographicai and bioclimatical characteristics Subsector lklt/Subbelt

all all

C.Mmdldo-Pw/aw/emm ~/a/cae Galaico-Portuguese

L)vnba/w/o- Trache//etum caende/ Galaieo-Portuguesr

Pob~0num c,~atum community Galaico-Portugtw.se

Mineashn

M/nc~-~ian

~ , Monuu~ M e u ~ ~ Supramed~crranean

Ti~mor

Thcrmo-, Eucollh~

T ~ r m o r

22 FOLIA GEOBOTANICA ET PHYTOTAXONOMICA, 28, 1993

Table 2 (com.) ~, , - -uni ty

Scctor Biogeographical and bioclimatical characteristics

Sebscctor Belt/Subbelt

G~lai,-o-Pomqptcse

Galaico-Asturian O~e~.4sp len /4mun conuuumse

Gaiaico-Portugtw~ Galaico-Ar~urian

Galaico-Porteguesc

Gahdco-,,e~stur~n Ses~m/do-Ananh/nemm duct/m/n/

Galaico-Porteguese

Oron ~n t)-Sanabrien ~i~ n

Galaico-Asturian saaa~ subsp. ~ community

Galaico-Astertan

Laeian0-Alacaren-~i~n

OY(~llf)ano-S9 n :~]~Cn ~;~ n

~ ~ w n gaya~ Ladano-Ancarendan OrensallO-~nahrie.n;ia n

Galaioo-Portuguese Lac~uo-,~m~rcr~ixn

Ore, nsano-S a n ~hrien~ n

sa~fra~ ~J~ae Orensano-S~n~hriensian

acro~ae Orensano-S~r~rienr ~-

O~'n~n,',-S*nal~cnsian

Orensano-Sanabrien~sia n

Orclkqal lo-S 8 n a b ~ n s a n

Sesamoido-Silenetwn gayanae

Orcnsano-Sanabrien~i~n

Mine~i:~n Compoaclan Northern Galaico-Asturian

M/neas/an Northern Gahico-Asturian

Mineasian Compo~elan Northern Galaico-Asturian

(~-cnsan

Northern Oalaico-Asturian

Naviano-Ancarensian Maragato-Stnthrien~i~n

Naviano-Ancaren~;~- Maragato.-Sanabr~nsian

Juresiaao~ueixensian Naviano-Ancaricnsian Maragato-S~n~hrien~i~n

Orcnr~n

Orensan

Orcn~n

Orcnsan Maragat o-Sanabricn~ia n

Orcnf.~n

M aragat o-S anabriendan

Maragato-S~n~hrien~ian

Maragato-Sanabrien~hn

Thermocolline Eucolline Eucolline

Thermo-, EucoUine Eucolline

Thermo-, Eu-, Uppercolline Thcrmo-, Eu-, Uppercolline Eli-, Uppercolline

Thcrmo-, Eucolline Thermocolline Mesomediterranean

EucoUine

Eucolline

Subalpine Oromediterranean

S~alpinc (h-omcd~crrancan

Uppermontanc Subalpine Oromediterranean

Mesomediterranean

Mcsomeditcrranean

Mesomediterranean

Mesomediterranean Supramediterranean

Mesomediterranean

Supramediterranean

Oromediterranean

Oromcditerranean

ORTIZ AND RODR~GUEZ-OUBI~: SYNOPSIS OF THE RUPICOLOUS VEGETATION 23

SYNTAXONOMIC SYNOPSIS

Asteretea tr/po///W~Trl, et BEEFr. in WF~'a. et al. 1962 Crithmo-A.nerietalia matitimae G~HU 1964

Crithmo-Armerion G~Hu 1964 Crithmo-Limonienion binervosi Gt~Hu e t G~nu-FRANCK 1984 1. Crithmo-Armerietum pubigerae ROZEIRA ex Plrcro DA SILVA e t TELES 1972

armet/etosum pub/geme J. et P. G u m ~ 1987 plantaginetosum mafiamae J. et P. GImaAN 1987 limonietosum binervosi P. et J. Gtmaku in ORTIZ e t RODRIGUEz-OumfiA (p. 30)

2. Spergulario-Armerietum depilatae FERUANDEZ PRm~ et LOIDI 1984 armer/etosum dep//atae FERNANDEZ PRIETO et Lotoi 1984 limonietosum binervosi FERNANDEZ PRmro e t Loml 1984

Asplenietea trichomanis (BR.-BL. in MEre et BR.-BL. 1934) OBERDORFER 1977 Potentilletalia caulescentis BR.-Bt. in BR.-BL. et JENNY 1926

Saxifragion trifurcato-canaliculatae RWAs-MARTII~Z ex R I V A S - M A R ~ IZCO et COSTA 1971

Petrocoptidenion grandifloro-viscosae FERNANDEZ ARECES, PENAS e t DIAZ- GoNZALEZ 1983 3. Saxifragetum trifurcatae ROTHMALER 1941 4. Petrocoptidetum grandiflorae LOSA, MAYOR, ANDR#_~ et NAVARRO 1975

Asplenietaliapetrarchae BR.-BL et MEIER in MEIER et BR.-BL. 1934 Asplenion petrarchae BR.-BL. et MEIER in MEIER et BR.-BL 1934

5.Asplenio-Cheilanthetum acrosticae SANTOS 1987 Androsacetalia vandellii BR.-BL in MEIER et BR.~BL 1934

Saxifragion willkommianae RIVAs-MARTIrCEZ 1963 6. Murbeckiello boryi-Sperguletum pourretii FERNANDEZ PRmTO corr. IZCO e t ORTm 1989 (p. 33)

sperguletosum pourretti FERNANDEZ PRIETO in ORTIZ et RODRIGUEz-OuBINA saxifragetosum spathularidis ORTIZ et RODP-dGUEz-OuBI~A

Cheilanthetalia maranto-maderensis SAENZ et RIVAS-MARTIrCEZ 1979 Phagnalo-Cheilanthion maderensis LOISEL 1970 COrr. I~REZ CARRO, DIAZ-GONZALEZ, FERSAUOEZ ARECES et SALVO 1989

7. Cheilantho-Asplenietum corunnensis PINTO DA SILVA COrr. ORTIZ et RODRIOtmZ- Otmt~A (p. 33)

Rumicetalia indurati Rrvas GODAY et RWAs-MARTtrmz ex RIVAS-MARTIr~Z, IZCO et COSTA 1973 Rumici-Dianthion lusitani RIVAS-MARTIrCEZ, IZCO et COSTA 1973

8.Phagnalo-Rumicetum indurati 17dVAS-M~TtNEZ ex NAVARRO et V ~ 1984 9. Sesamoido-Anarrhinetum dwimini OR'nZ et RODRIGUEz-Otmn~A (p. 36)

ana~/netosum dudmin/ORTIZ et RODRIGUEZ-OUBI~A rumicetosum indurati ORTIZ et RODRIGUEz-OuBI~A

Saxifragion continentalis RIVAS-MARTINEZ ex RIVAS-MARTINEZ, FERNANDEZ GONZALEZ et S,$d'/CHEZ-MATA 1986

10. Phalacrocarpo-Saxifragetum continentalis ORTIZ et Izco in PI~REZ CARRO, DIAz-GONZ.4a.EZ, F. ARECES et SALVO 1989 (p. 37)

Cheilanthion hispanicae RrvAs GODAY in RIVAS GODAY et al. 1955 era. SA~rZ et RrvAS-MAR~Z 1979

11. Linan'o -Cheilanthetum tinaei FERNA2,rDEZ ARECES, P~REZ CARRO et

24 FOLIA OEOBOTANICA L~ PHYrOTAXONOMICA, 28, 1993

DIAz-Go~rz~aJ~z 1987 �9 che//amhetosum tinae/Fm~4a, rDEZ ~ Pta~z CA~O et DIAz-GoNzt~z 1987 che//anthe~osum h/spardcae FERNANDEZ ARECES, P~tEZ CARRO et DIAz-

Gorcz~ .~ 1987 che//ant/~osum 8zamch/cae ~ E Z ~ P~P.Ez CARRO et

DL~Z-GONZ~L~7. 1987 Anomodonto-Polypodietalia O. DE BOL(~)S et VivEs in O. DE B o t ~ 1957

Bammnio-Polypodion cwnbrici O. DE B o i l s et V~ms in O. DE BOL~ 1957 12. C ~ D a v a / / / e t u m canar/ens/s B~LOT et ~ in ~ E C A 1959 (p. 4O)

dava///eto~an canar/ot~ B~.LOT et ~ in OgTm et RODRIOUEz-Oumg~ po~'pod/~oslor/ /nter]ect /ORTIZ et RODRIGUEz-OuBI/~A

Padetadeta//a R r V A S - ~ ex RIVAS-GODAY 1964 Parietario-Galion murale R I V A S - ~ ex RIVAS--GODAY 1964

13. Par/etar/emm juda/cae AR~T~ 1928 14. Cymba/ar/o-Trache//etum caendei RIVAS-MARTINEZ 1969

Polygomon capitatum community (p. 42) Th/asp/efea rotund/fo///BR.-BL. 1948

A ~ a / i a a/pmae BR.-BI. in BR.-BL et JE~qY 1926 Linado-Senecionion carpentan/RIVAS-MART~Z 1963

Rwn~ scutams L subsp, ga//aec/cus community (p. 43) 15. Cryptogmnmu~Silenetwn gayanae FERNANDEZ PRma~ corr. ORTIZ et RODRIGtmz-OueI~. (p. 45)

Rumex suffrut/cosus community (p. 45) 16. Sesamoido-Silenetwn gayanae Izco et ORTIZ COrr. Omaz et RODRIGtW_z-OtrBI~ (p. 46)

s//enetoswn gayanae Izco et ORTIZ in ORaaZ et RODR/Gtmz-OUm~A /ep/d/etosum reverchon//ORTm et RODRIGUEZ-OuBII~A

Po~stichetalia lonchitis PdVAs-MARTI~Z, DIAZ-GONZs FERNkNDEZ ~ , LOIDI et PmcAs 1984

Dryopteridion abbrev/atae RIVAS-MARTtNEZ 1977 17. Ctyptogrammo-Dryopteridetum abbreviatae RIVAs-MARTINEZ 1977

DESCRII~ON OFTHE NEW SYNTAXA AND COMMENTS TO THE SYNTAXONOMIC SYNOPSIS

The synoptic table (Tab. 3), representing 185 relev6s, summarizes the floristic composition of the associations of the Galician rupicolous communities. It can be seen that, in general, the differences between the characteristic species combinations of the associations included in the synopsis are sufficient to be of diagnostic value. Equally well defined are the three phytosociological classes included in the synopsis: the rocky shore halophytic communities of Asteretea tr/po///, and the non-littoral communities of Asp/en/etea tr/choman/s (on consolidated substrates) and Thlaspietea rotundifolii (on mobile substrates). Within Asplenietea trichomanis there is clear floristic divergence between the calcicolous associations of the order Potentilletalia cau/e.scent/s and the remaining orders of the class. In the case of these latter it is apparent that the taxa normally considered as characteristic do not serve to differentiate dearly between them. With reference to the syntaxonomic synopsis we will here comment only upon new syntaxa, proposed corrections to existing syntaxa and syntaxa which have not yet been comprehensively described.

oNrlZAND R O D P . / ~ ~ OFTHB RUPICOLOI~ VI~JErATION

Tsbk 3. Syuot~ tsbZc

Set o f r c l c ~ Numbs" of spa~cs

per column Number ~ rclcv~

1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17

23 14 29 31 13 22 20 49 54 38 29 39 9 10 23 43 24 6 7 5 16 8 10 5 9 15 11 20 27 2 3 2 15 9

Dis~p~,zic tam ot msociation

s u t ~ . ~ z m E

~ mm~na sm~n~ ~

s u b s p . ~

Vat . C~NOIn4~L~

N o d s o ~ mm~at~

J ~ m ~ o n ~ o l ~ n Ammb~uon dsm~s~m

Anm#smzun x bsZemw.d~zm

Ose//ms0~ x ~ c a ~ x~rulm~

Dm~l~ cmsm4zn~

T ~ c~md~von

S ~ a e f o e ~ subsp. Sesmno~e~ m/~r

ii

n l

V

in n I

I

H

I H

I

IV

�9 I n I I

26 FOLLA OEOBOTA,'CtCA El" PttYTOTAXONOMICA, 28, 1993

Set of relev~ 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17

Tua o fA .m~a o~oo~

Fe.ma~ mbm

sut, s p . ~ m v s, oap,/a~ a,,o~la

Tan ofAw/,m/e~ O/choman~

Pha~a /on ,~,a~_!., A.vp~na,m v~oman~ ~ o~aan ,m

s e ~ m ~ Fgus c . w ~

T a m c/Potent/Heta~ caule.w.en~

~ fa~mus Cam, oanu~ a , v a ~

sul~p. ~ Mau~o/a _fr,~_'_ __!osa

~ o ~ , n .

subsp, amnwa/di/

ra~p. ~nmata

subsp, a.n~ca ruta..nnow~

P e ~ c o ~ ~ c o s a

subsp, t e ~ , ~ m

Se.dum k~-smon

H V V Ill I IV II i n

II I II V

11

im i v n V IV H

IV V I n

IV H H IV II IT

I I

I I

[ ] H

H IV

H I

V I . Il l I �9 I H HI

I II

I I

HI IV

I

V IV II IV

1I H HI I IV I III m III

V

Olgrlz A . ~ RODRIGUEZ-OUBII'TIA: SYNOi~S OF THE IRUIqC.OLOUS VEGETATION' 27

Set of rek 'v~ 1 2 3 4 5 6 7 8 9 10 11 12 13 14 1.5 16 17

T a n of Rte,,~e~_~_'e/nduma

ttdmp. ~ o ~ m t It II I I

~ e a e m I

C ~ t u ~ m a ~ ~

T m o i ~ ~ o ~

~ t / f l ~ m / m m ~ I . I V V m m v

O~,fa~o~n m

O ~ m a m ~uanem~ I

O t l ~ t a m

~ l ~ f f o ~ I . IV H H IV IV II

. ~ . ~ n m m n i i ~ z t / s puOm~.a I m I L M ~ m . m IV II I I

tulmp, dwfeu/ I I I I

Y ~ m iI H IV I I R u m ~ a~fo~e//a u . I . IV I I

A ~ m ~ a t ~ II ~ , ~ , a , , _ , . I I

subsp. ,Z~z~ X I m ~ ~ I I I ~ e/atiua I I II n ~ M , ~ , ~ x n H

I HI V

IV

�9 I ] ~

V m I

m i

I m I I

�9 IV

V v I m

m I

28 FOLIA OEO~OTAN~C.A L~r ~P/'rOTAXONOM~C-.A, 28, 1993

Set of reicv~ 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17

II 1I I �9 HI Ill I

sanf,~ ~

z ~ o n p y ~ c u s

z~aa c a e ~ Fesmca d . r/m/av/a F e a m c a / n d g r ~ S/Zem a u t 4 ~ s.a.

..ge.dum ~ z a m ~ a

Nm'c/ssm ~ lubsp.

Fe. .m~ d lbe3,~ Ym'/oae/aev/s

s a b 6 p . ~

e z ~ c.ormo~ $ / ~ m u a # h m

Yamkzam/r Lcoatodon ~ H

S o a O ~ o/emcm., I

~ , a o a ~ ~ ka:/dum Gema/zon ~

~ e . a m ~ u m

A,~b~ ~ o ~ L o c a ~ ~ a e a

~ m ~ m ~oaaum F e . m ~ e.~an.v

Hon/avJa scabtfda

5//erie a~a

Hedem he/ix

R~eda med/a

Rubus sp.

Mocut/a/is aetna

HI HI IT I

H I I U

H U I

IH I

I HI

HI

I I I I . HI I

I I

I

H

II

I m [ ] H I I I I I I II

I I

H I II I I I

HI

I

I I

I !1

HI II V I

HI I II I

V I I

V I I

m i v

Ill

H I

ORTIZ AND RODI~GUEZ-OuBI~: SYNOPSIS OFTHE RUPICOLOUS VEGETATION 29

Set of re levis 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17

c ,~o~y~a I H G a ~ ~ c y n ~ m

subsp. ~/v/anum V I p/anta~ a~na IH I ~/mm athanum~:um IH I

I~ cee column Column 1: Aspleniwn madnum II, Atnplex hastata I, Daphne ~ i , ff,_,_m I, Inu/a ~ I, lunoa nut, e/mus I, goelaia glauca If, Plantago lanceolata I, Scirpus holoschoenus I, Trifolium occidentale I. Column 2: Daucus caroca subsp, gumm/fer I, Euphorbia portlandica I, Limonium biner~osum I, Runu~ acetosa

cob- - . 3: An~none palma~ I, A,~/s athena I, Cynosuna ~ L Col-ran 4: Anner/a sp. III, Artenus/a cwnpes0~ subsp, glutinosa HI, Aspemla cynanchi~ I, Dianthus hyssopifolius i, GaUum m u , ~ H, thymus zy~ m. Column 6: H/am'bins pa///dum subsp. ~ran/~zum I, Phyceuma hem/sphaer/cum I, Poa/e~onens/s I. Column 7: Brachypodium nq~es~e I, Heraclewn sphondylium I, Leucanthemum g ~ , _ ! ! ~ I, LoDem gallica I, S//ene v u / ~ I, Th/asp/a/pe.n~ subsp, met/no/IL Cohmm 8: ~ spTumvcepha/on I, Awna barbara H, B~a ~ II, Campanu/a h~w~ca I, Ca~_ l,,- carpetanus I, Cenvunthus calc/~apae II, ~ ech/,#~, I, C3~nosun~ degans I, Daucm caro~ subsp, carom II, De.tmaze~ rigida I, Foeniculum vulgate II, Lactuca seniola I, Lavandula stoechas subsp, sampa/ana HI, Pe~vhagia nanteuilii n, Suta montana I, San~orba minor subsp, mag, u ~ 11, Santo//na r o s m ~ o / / a subsp. sem/dentata I, V'w/a sat/va H, Thymus mast/ch/na H. Column 9: Adenocarpus comp//cams subsp, complicatus I, Adenocarpus c o m t a i ~ _ subsp./a/nz//I, meonanthum subsp, salcedoi I, Centaurea aristata subsp, langeana I, Centaurea m/crantha IL Co/ncya sp. I, Cy6.n~

I, Echium rox,~!_r4,__~m I, Evica cinerea I, Galium apa:ine I, Galium rivulare I, Hyperirum iinarifoliwn H, Hypocho~ radicata II, Lapsana communis I, Rumex acetosa subsp, acetosa I, To/pis ba~ata I, Lm, andu/a ~echas subsp./uis/en I, P s e u d a m ~ e ~ longifolium I, Tuberaria globuladfolia I, Colunm 10:. Anthoxanthum odoramm l, Arenar/a montana I, Armer/a grex m~t~mntana I, Asp/en/um onopte~ I, Gagea pmten~ I, L/thodora d/ffusa I, Ompha/odes n/t/da I, Poa bu~osa I, Sag/fragu c/us//subsp./ep/rm/h~ma H,

Column 11: Fumavm rnura//s I, Osmunda rega/~ I, Osy~ a/ha I, PIe6d/mn aqu///num I, Sedum se.~-fom~ I. Column 12: A//fitm car/hahn I, Ca//una vu/~/s I, Centaurea pan/o,C~_o I, ~ c/av~_ ,~ I, Er/genm ka~insManus I, Gagea nevaden~ I, Lithodom prostrata I, Omi~/topus pcpusillus I, Ranunculus pahutmu~ I, Seneci.o sylvazict~ I, Senecio vutgads I, Tee.utalia nudicaulis ~ Tube.rana Kuttata I, U~r.a memb~,w~:ea L Column 13: Holcus ianatus H, Poa annua I, Urtica dioica I. Column 14: ~ cap///ar/s II, S a g ~ n a ~ b e n s V. Column 15: M ~ sedo/des HI, S/~me ~ HI, Sol/dago ~,a-aurea HI, T~/fo~um alp/hum m. Column 16: A~ost/s sto/on/fera I, ~ minus I, Co/ncya pseudoemcasm~ subsp, s~sera I, C3~zus omme__~!e, nmeus I, Hieracium pilosella I, Jasione crispa subsp, brev/sepa/a H, Leucanthemops/s pqj/'v~_ subsp. flaveo/a HI, Nardus s~ffa I, Poa pratens/~ L Sedum an~//cum subsp, pyrena/cum I~ Spe~u/a ~ I, Teesdaliop~ conferta I, Thymelaea dendrobryum I. Column 17: ~ lampsanoides I, Epiiobium angusu'folium I, G e ~ n a &tea I, Juniperus communis subsp, a/p/ha II, Senecio nebn~nsis I, Succisa pratensis I, Vaccinium ~ H, V'uTla ~ I.

30 FOHA OF..OBOTANICA El" PHYrOTAXONOMICA, 28, 1993

Reference of the association tables: Cohnnn 1: C-Athnm-Annoietum pubigerae, table 4 in this article. Cohnnn 2: $ ~ A n n e r / e t w n dep//atae (FERNANDEZ PRIETO and LOIDI 1984, table 1: relev~s 1, 2 and 16), (1'. Gum.AN ~989, table 23). Column 3: Sa~fraSeaon ~fu,,r.atae ~ 7 - A R E C E S et al. 1983, table 7). Column 4: Parocop6delum ~mdiflorae 0..OSA et aL 1975, table 2). Column 5: " " acrost/cae (FERNANDEZ-~ et aL 1987, table 3). Column 6: Mwbeck~lio-Spogdaum poutretii, table 5 in this article. Column 7: Ou~//antho-Asp/en/emm comnnense, 5 unpublished relev~. Column 8: ~ R u m i c e z u m /ndur~ (ORTIZ 1986, table 27), (FERNANDEZ-AREr et al. 1987, table 2: relev~ 2 and 3). CoJumn 9:. S~amo~o-Anat#u.tun~n dw/m/n/, table 6 in this article. Column 10: Pha/acrocarpo-Sa6fra~um cont/nenta~ (ORTIZ 1986, table 19), ~ - C A R R O et aL 1989, table 7: relev~ 4, 5 and 6). column 11: " " " 6nae/(FERNANDEZ-ARECES et aL 1987, table 1: relev6s 1-15 and 17-21). Cohnnn 12: G y m n o s r w n m o - ~ canat /ot~, table 7 in this article. column 13: Par/etaraum j u , ~ " ae (ORTIZ 1986, tables 28 and 29). Cohnnn 14: C)vnba/a~7-Tmche/~um caou/e/(RIVAS-MARTINEZ 1969, table 3). Column 15: ~ $ ~ n e t w n gayanae (ORX~Z 1986, table 23). Column 16: S e s w n o / d o - S ~ gayome (ORTIZ 1986 table 24: relev6s 1-3), ~ and ORTIZ 1987, table 3), table 11: reJev~s 1 and 2 in this article. Column 17: ~ D ~ p t e t / d e m m abbre~_'~ (ORTIZ 1986, table 21), (Izco and ORTIZ 1987, table 1)

1. Crithm~Annerietwn pubigerae ROZ~mA ex ~ DA SILVA et ~ 1972 Table 4 atmer/etomm pub/gerae Table 4: relev~s 1-5

Ilmonletosum binervosi P. et J. G u m ~ in OR'nz et RODS/GUEz-OuB~A subass, nova hoc loco Table 4:relev6 6 Holotypus: table 4:relev6 6

The association Crithmo-Annetietum pubigerae is found on the cliff~ of the whole western side of Galicia and a large part of the northern coast, areas which have mainly gneissic-granite substratum. However, in the small quartzitic outcrops which occur within its area of distribution, it is differentiated by the presence of Limonium binervosum (G.E. SM.) SAX~ON, thUS defining a subassociation limited to this type of rock (P. G t n T t ~ 1989). Quartz becomes dominant in the extreme Northeast; on slate substrates, the other dominant rock of the Northeast, this association is replaced by the Spergulario-Annetiemm depilatae.

ORTIZ AND RODRIOUEZ-OUBI~ SYNOPSIS OF THE RUPICOLOUS VEOETATION 3]

Table 4. ~ A m t o / e a o n p u b / ~ RO2~-RA ex PINTO DA SILVA CA Tl~ ~'~ 1972

- sabass, wmo/aosum pubi~me J. et P. GUITIAN 1987 (rclev~ 1-5)

- subass.//mon/aoaon b/n~vo6 P. et J. GurrIAN in OR'I1Z et RODRIGUEZ..OuBI~ (rele~ 6)

Relc~ nnmber 1 2 3 4 5 6

Altitude (m s.m.) 10 2O 10 20 10 10 Orientafi~ NW W SW W SW SW b ~ e i o a (o) 6o 6o .so 70 7o 80 Area analysed (m 2) 10 25 25 50 25 10 c o ~ of v ~ a ~ . ~ (o~) 6o 45 25 40 4o 2o A ver~ height of vegc~ion (an) 20 30 20 25 30 35 Nmnber of ~ecies pet relcv6 5 5 6 7 8 6

Di~noaic tam of at~adafion

Anng/a pu/,ip~a subsp, p~igca

D~snost~ tam of J,n;~n,~ and cfder

2.3 33 1~ 33 1.2 L2

Cr/t~num ~ 3.3 2.2 2.2 1.2 F~tuca mbra ~p. pruino~a +=2 Z~/.~ mar/aa +

SpeCtra ,u,0~a +

Diagnr ta,~ of class

Differem~l tan of subass.//mm/e~osum b/nen, os/

L/mort/urn b m

Olhcr rain

xogo~ ~ P~au~so ~ Sonchus o/emceus

S~ene un/fo~ ~ a ~ c ~ re,v/area

SC~Xa ~gosOmema Carana ~ a

1.1 + + + +

+ L1

+.2 +

+.2

2.2 +

+

+.2

+.2

1.2 1.2

1.2

32 FOL/A O~OBOTA~C.~ L~ PHYTOTAXONOMIC.~ 28, 1993

Table 5..Mwbeck/e~ bo~-Spe,jpdeaun p o u m ~ ~ e Z PemTO corr. Izco et OR'nz 1989 (* - no data provided ia the r, o e r ~ pebticat im)

- sltbass. ~u/4~toe~n pcmme~ PERNANDEZ PRIETO in ORTIZ et RODItlGUEZ-O~ (relev~ 1-7) - subau. ~ #pathu]a6d~ ORTIZ et RODItJGUEZ~ (relev~ 8-10)

Ite.le~ m m l ~ Akitede (m ..~) Orientl tm lw l i , , t i ~ (o)

Area analysed (m~ Cow.r ot w'.mlar ~. . t , (%) A ~ a ~ h~ht ot ~ l r . t , ~ (m) Numb~ of rock.s per

1 2 3 4 5 6 7 8 9 10 2100 2100 2010 2010 2080 2050 2000 2050 2070 1940

N N NE NE E SE SE NE NE E 90 80 80 90 -* - 90 80 90 20 10 30 20 10 10 20 10 10 20 5 10 5 5 - I0 20 5 8 8 10 10 - 15 5 5

11 7 8 6 6 5 6 8 7 5

D i ~ ~ of auor and InliM~

Diasaeak m a of dms

Sedum Ignua#n

Di~ert-mlal taxa of

S a ~ ~pathu/a~

Other tala

Se~n bz~#'o~n

Fe.nuca IJu~na s.l. A y o ~ akn~

+.2 1.1 L1 1.1 +.2 2.2 L2 1.2 1.1 +.2 L1 2.3 2.2 1.2

L2 +2 2.2 1.1 L2

2.2 2.3 +.2

1.2 +2 + +.2 +.2 +.2 +.3 1.2 1.2 1.2 1.2 + 1.1 +.2 1.1 1.1 + + 1.1 +.2 r +2. + +.2 +.2 +

L1 + +.2 + +2. r 1.2

1.2 + 2 + +'7

Tram present in only oae rele~: A ~ o ~ c.ap//Z~b +, in Relev~ 1; A M o ~ tnmc_,~_~!_~ subsp, dw/eu/ +, in R. 3; ~ oreades +2, in R. 7; H'u~umsm~/~zmm +, in R. 10;,H'u~rump~_/_/d~_~m subsp .~m~/~n 1.l, in R. 7;Luzu~ cae.sp/tr r, in R. 9; Phy~uma hem/.vphaeHcum +.2, in R. 1;eoa/e~mens/s +, inR. 9 ; , P ~ ~ +.2, inR. 5.

OErlZ AND R O D I ~ G ~ U B ~ SYNOPSIS OF THE RUPICOLOUS VEGETATION 33

6. MurbeckieUo b o r y i - S ~ powretii FERNANDEZ PRm~ corr. D.CO et OR'nZ 1989 Table 5

~.rguletosum pourretil FERNANDEZ PRIErO in ORTm et RODRfGUEZ-OUBI~ subass, n o v a

hoe loeo (FeRrCg'~EZ Psmro 1983: 497, table 5: relev6s 1-3; relev6s 5-7 in table 5 of this article). Table 5: relev6s 1-7 Holotypus: the same as that of the association: table 5: relev~ 2 ( ~ E z PtmTO, 1983: 497)

�9 saxttl'agetosmn spathularldis ORTIZ et RODRIGtmz-Ou-eI~ SUhaSS. n o v a hoc loeo Table 5: relev~s 8-10 Holotypus: table 5:relev6 9

The communities described as Cryptogrammo-Saxifragetum spathularidis F. PRIeTO, J. G u m ~ et AM1c, o 1986 from similar habitats in the subalpine belt of the Serra dos Ancares, appear to correspond to very impoverished units of Murbeckiello-Spergulaum pourretii sax/fragetosum spathu/mid/s. Two taxa which characterize this association, and which give i t i ts name, are not present in the Cryptogrammo-Saxifragetwn spath,_d_midis, so we reject the previously proposed poss~ility (ORTIZ op.cit.) of combining it at the subassociation level with Murbeckiello- Spergdarw: powrefii.

It shows affinities with the endemic association Saxifrago-Murbeckielleatm henninii Bit.- Bt., P. SILVA, ROZEmA et Forrms corr. RIVAS-MARTIiqBZ 1981 of the Serra da Estrela (Portugal), characterized both by its partially endemic floristic composition and by another subspecies of MurbeckieUa boryi: M. boryi (BoISS)." Ro'rriM. subsp, herrmh//RIVAS-MARTIrCEZ, whose populations, although showing some deviation from the type subspecies, have not been considered sufficiently differentiated by some authors (FmmAm)ES 1950) to have taxonomic independence.

The new subassoeiatiom are found in the Oromediterranean or Subalpine belts of the high siliceous mountains of the Orensano-Sanabriensian sector and east of the Orocantabrian Province.

The second subassoc/ation develops in very humid fissures and on ledges of the quartz and slate crags situated above 1,900 m in the Macizo de Pena Trevinca, placed in the Oromediterranean belt. It is distinguished from the other suhassodation which occurs in drier localities, by the presence of Saxifraga spathularis BROT.

7. Cheilantho-Asplenietum conmnensis PINTO DA SILVA corr. ORTIZ et RODR[GUEZ-OUBIIT/A hoc Ioco (PINTO DA SILVA 1965: 17, synoptic table)

Following Article 43 of the Code of the Phytosociological Nomenclature, we hereby correct the name of the described association of the serpentines of the northeastern Transmontano area of Portugal originally named Cheilantho-Asplenietum cuneifolii by PINTO DA SILVA (1965). Correction is necessary because the representative of the genus Asp/en/um is in fact A. ad/antwn-n/gnan L. var. corunnense ~ in accordance with NOGUmRA and ORMOtCOE (1986) and HORJALF_S (1986). To this association we therefore classify (at least

34 Fot,~ OEO~OTAN~CA In" ~FrrOTAXONOM~C~ 28,

Table 6. S e a ~ m ~ / m ~ ~ ORTIZ r RODItlO~

- mbua amm,~m~eum ~k##~ Ox'nz et ROD~GUEZ-OUm~ (relev~ Z-8) - mbaf~ n#nir.aos~m ~ Ot'nZ et RODglGUEZ-OUm~ (relev~ 9-15)

ReJe~ number 1 2 3 4 5 Akitede (m s.m.) 2"/0 90 200 110 400 OrL-~-t=~- SW SE W SW E |nel;n~nn (0) 2 95 1(}0 80 gO A~ea ~/u,d (mb 3 m 10 5 10

Cover of v .m~r l ~ m (%) ~0 45 40 9o 5o AveraSe h e ~ of vegetadm (cm) 20 45 40 50 50 Number of ~eie~ per relev~ 6 11 S 17 12

m:m d ~ and .m. , ,~

~ x #u~mmt~m Meac4 ds~e mbtp.

]~/~-~-~.'~.l t a n of ~ mm_~-_ _ _ ~ m

Other uum

/ h m ~ ~ a a

~ m

1.1 1.1 1.1 §

2.3 +

L1

L1

2.2

L1

2.2 2.2 +

+.2

1.2 §

+.2 4-

2.2 + 4- + +

1.2 I.I 1.2 + + LI +

4- § +

§ r

LI + L1 +.2

+ + L1

+

Ogmz AND RootkRr~-O~ S~wo~s Ol~ Tim RtnqCOLOUS VBOEr^~ON 35

6 7 8 9 10 11 12 13 14 15 480 340 400 420 150 400 80 90 280 580

S E E W N S S S E NE 75 90 75 100 100 95 100 90 95 75

7 10 5 10 15 10 10 10 10 5

40 65 70 85 30 85 75 75 70 60 40 70 60 50 50 50 70 60 70 50 13 17 15 II 17 22 19 22 20 17

L1 2.2 L1 2 2 L1 L1 Z2

+.2 + 1.1 L1 +.2 +

+.2 +.2 r

L1 L1 +.2

L1

r

r

1.1 +

+

+

§

+ + + + + L1 + + +.2 + +

2.2 +

+ ~

r

.2+ §

L1

1.1

3 3 2.2 3.3 1.1 L1 +

2 3 2 2

+ 2

12

+ + 1.2 +

12 + L2 +.2

r + r L1 + +

r

§

+.2

+

§

r r

f

+.2

+ +

+ + + +

+ 4- r

4- 4-

L1 r +

+ r

L1 +.2 +.2 r

+.2 +

r

1.2 + +

§ r

§

+

r +

.f

+.2

+.2

36 FOL~ oeoeo'rAmcA e'r ruv'rorxxoNoe, ac,% 28, 1993

R e k ~ number 1 2 3 4 5

Ceatma~ ntknmt~

$//r162 autam ta . To~ l ,~aa

Lacaw~ v/m/nea

4-

+

4-

T m la~eat in oaly oae or two re.lev~ Adenocw/aa ~ ~h~_~./a/nz//r, in Relev610;, r, in R.LS. ~ ~ subsp./aa~.ana+, in IL6; +, ia R . 1 4 . ~ j a m c . e a +,inlLg; +,inR.12. Ga//wn ~ +.2, in R.11; + , in R.12. Rum~acefo~ r, in R.4; r, in R.11; Sedum fo~enmum +.2, in R.9; 1.1, in R.11. Adenoe.a~s ~ subsp, compi/ca~ r, in R.14. A~m.~ ~uncaZu/a sulatp. ,~w/eu/+, in R.6. ~ / r +, in R.8. C)~/.ms mu~7om8 r, in t .7. F-,ch/um

provisionally) the rupicolous communities of consolidated rocks occurring in serpentine areas of Galicia, although the intricate nomenclature problems posed by this type of syntaxa described by PINTO DX SmvA (Op.CiL) call for a more detailed stucly. This will be the subject of another paper.

9. Sesamoido-Amuchiaetnm durimini Og'r~ et RODRIGUEZ-OUBII~IA aSS. nova hoc loco Table 6 Holotypus: table 6:relev6 8

anarrikinetonm durimini ORTIZ et RODRIGUEZ-OUBII~ subass, nova hoc loco Table 6: relev~s 1-8 HolotYIa~: the same as that of the association: table 6: relev~ 8

rumleetosma iaduratl OR'nZ et RODglGUEz-Oue~A subass, nova hoc loco Table 6: relev~s 9-15 Holotypus: table 6:relev6 14

This association generally colonizes natural or artificial stony slopes. Since/t tolerates high-nitrogen conditions, it is frequent on roadside banks.

It is distributed mainly in the coiline bioclimatic belt, above aU in the ThermocoU/ne Subbelt, and particularly in the lower levels of the Upperconine SubbelL

The new association is very reliably characterized by the presence of the two taxa that give it its name: Seswno/~ suJj~ut/cosa (I..~GE) O. Kumze and ~ dur/m/num (BRO'r.)

both endemic m the northwest Peningula. According to ~ z RODXlGL~Z et al. (1989), towards the Mesomediterranean belt of the I.~sitano-Duriensian sector there occurs a community strongly related to the one des~bed here, called /man/dno

OltTlg AND RODPJGUEZ-O~ SYNOPSIS OF THE RUIqCOLOUS VEGETATION 37

6 7 8 9 10 11 12 13 14 15

+ 1.1

1.1

1.2 +.2 +

§

1.2 + + +

farm/atom r, in R.12. E6ca c/naea r, in R.15. Ga//mn a ~ r, in R.13. Hedua he.h~ + .2, in R.13. Hype6cum p~ommm +2, in IL13. Lapsana commun/s +2, in IL12. ~ sunr.~ subsp. ~_C'_~ r, in IL13. Nan~aus u/angus suhtp. ~/andms r, in R.10. Pw/aw/a ] , , ~ r, in R.11. P s ~ /on~o//um +, in R.2. Raeda med/a +.2, in R.8.Rubus sp. +.2, in R.& 7k/~rar/a~obu/at~o~a r, in ILg.

dw/m/n/-Rum/cetum/ndurat/. In this community, however, Sesamo/des s u ~ (LANGE) O. KUN'rZE does not occur and Rumex/ru/uratus BoLss. et REUTER is a constant species. Two additional taxa, both absent from the Galician association, D/anthus ~ t a n u s BRor. and Ant/rib/hum gran/t/cum ROTHM, also occur there. This association is probably a Mediterranean vicarious association of the S e s m m ~ / d o - ~ dur/m~, which has an almost entirely Eurosiberian distribution in Galicia. The other associations of the alliance RumiciDkmth/on &.v/tan/have a floristic composition very different from the Ananh/nenon dut /m~.

At the boundary between Mediterranean and Eurosiberian regions, specifically towards the Mesomediterranean belt of the Orensano-Sanabriensian sector, Rum~/nduratus BolSS. et Reuteat and Erys/mum//n/fo//wn (PEgs.) GAY subsp. ] J ~ f o / ~ become incorporated into the association. These two species characterize the subassochRion S e s a m o / d o - ~ dur/mbtf r i C o . c u r e /ad/Ara~, which is typical of this tran~itionaJ zone. Inside the Orensano-Sanabriensian sector, it is replaced by the ehagna/o-Rwn/cetum/ndurat/, with the endemic taxon Ananh/num due/m/hum (BgOT.) PERS. being replaced by A. be///d~o//wn (L.) When. Probably, towards the Lusitano-Duriensian sector, this same subassociation marks the tral~tion towards the community mentioned by Ss RODRIOU~ et aL (o13. cit.). From the biogeographical point of view, the new association is distributed throughout the Galaico-Portuguese sector and penetrates slightly into the western extreme of the Orensano-Sanabriensian sector.

10. P/~_l__,~'rocarpo-Sa6f~emm cont/nenta//~ Ogrlz et Izco in ~REZ CARGO, DO, z-Gotcz~I ~7~ ~ E Z A g e c ~ et SALVO 1989

Stands (not sampled in the present study) closely related to this association occur in the Supramediterranean belt of the Maragato-Sanabriensian and Transmontane subsectors and

38 FOLtA GEOBOTANICA ET FHYTOTAXONOMICA, 28, 1993

Table 7. Cowmo/Fummo-Dam///etum c.anat/en~ BELLOT et CASASECA in CASASECA 1959

- s u b a s s . ~ / m e r j e c t / B E U . O T et CASASECA in ORTtZ et RODRIOUEZ-OUBI~ (relev6~ 1-17) - subass, dava//ietatum canar/ensf$ ORTIZ et RODPJGUEZ-OUBINA (relev~s 18-27)

Relev~ number Altitude (m sJn.) Or ien t a t iOn

Itw]ina~Ola (o)

Cover of vascular plantg (%) Average height of vegetation (cm) Number of species per relev6

1 2 3 4 5 6 7 8 9 10 10 530 220 10 10 40 240 240 10 80 E SE W W SE W NE NW E N

90 90 90 90 90 90 80 90 80 70 2 2 1 3 2 1 4 3 2 0-5

25 20 30 30 20 40 60 60 25 20 20 15 20 20 25 20 25 15 15 15

8 6 7 9 8 7 7 8 8 5

Diagnostic taxa of ~ t i o n and allianCe

Pob~oaum ~,:oje~. Umb~:us .qw.s~[s

Dam///a

Diagnostic ta.xa of order and r I ~

Sedum hi, suture

Other taxa

Hedera ~./~ s~sn b,r~'ot~on Narc/ams tdandms

Se.dum arenanum ]as/one montana Un/ca membmnacea

P, amozu~/~_ ,a,~sus ~ p m s m u a ~ , n u n ~ Se,m/o Pa~.tar/a ]uda/ca

Tuberwia ~_:~t~_ Reseda media Silene nutans subsp. Linaria saxa~lis

S e ~ g l ~ w n

2.2 3.3 3.3 33 2.2 3.3 4.5 3.4 2.2 1.1 1.2 + 1.1 12 + 22 12 2.3 + 1.2

+ + i . i + 1 2 + +

+ 2 + + + r + 12 + 2 1.2 + + .2 + .2 + .2 + + + 2 + 2 1.1 + 1.1

+ 2 + 2 + r + 2

r

+.2 +

r ,

r

+=2

r

+ 2

O E r l Z AND R O D I ~ G U I ~ - O U B I ~ SYNOPSIS OF THE RUIqCOLOUS VEGETATION 3 9

11 12 13 14 15 16 17. 18 19 20 21 22 23 24 25 26 27 50 240 340 450 450 460 480 120 110 30 10 80 100 290 360 310 398

SB NE NW S W N W NW W W S N NE W N NW S 90 60 90 90 85 90 90 80 70 60 90 80

1 3 0.5 - - 2 2 3 1 1 4 1 1 0.5

30 20 15 - - 30 25 30 25 20 25 15 25 20 20 15 25 - - 25 25 25 30 30 30 25 35 30

9 3 2 9 12 10 8 7 9 9 6 7 8 8 3 3 10

L1 2.3 1.2 3.3 + + + L 1 + .2 1.1 1.2 1.1 L 1 + .2 + + .2 +

L1 2.3 1.1 1.1 + 1.1 + + + 1.1 r + 2 L 1 + r +

+ L 1 + + + + + L 1 + +

2.2 2.2 1.2 1.2 1.1 1.2 1.1 1.1 1.1 +

1.2 + + + + + 2.2 + 2.2 + .2 + +

+ + + + + + .2 + + .2 + + + + + + + + +

r + + 2 + 2 r + 2 + + + §

4. + §

§ + +

+ +

+ +

+ +

r

+

+.2

+.2 4-

§ r

40 FOLIA GEOBOTANICA El" PHYTOTAXONOMICA~ 28, 1993

Taxa present in only one relev~ (Cont. Tab. 7.): Centaurea paniculata +, in Relev6 27; Chelidonium majus r, in R. 15;. Engeron kwv/nsManus +, in R. 1; Gagea nevadensis +, in R. 27; Omithopusperpusillus r, in R. 5; Phagnalon saxat//e +, in R. 8; Senecio vulgwis r, in R. 11; Silene a/ha r, in R. 16; Sonchus o/r r, in R. 6; Trachelium caendeum r, in R. 8.

in the Montane belt of the Juresiano-Queixensian sector. The floristic composition of these stands is very similar to Phalacrocarpo-Saxifragetum continentalis, but the Phalacrocarpum is represented by P. oppositifolium (BROr.) WILLr~ subsp, hoffmannseggii (SAMP.) NmTO instead of the typical subspecies, whilst Dianthus lusitanus BROr. appears frequently. These stands, which border the distribution area of Phalacrocarpo oppositifolii-Saxifragetum continentalis on the south, may represent a subassociation of the latter or an independent association. The data currently available do not allow us to incline, for the moment, towards any one of the two options.

12. Gymnogrammo-Davallietum canariensis BELLOT et CASASECA in CASASECA 1959 Table 7 Lectotypus: table 1:relev6 4 (CAs~ECA 1959: 325)

davalUetosum canariensis BELLOT et CASASECA in ORTIZ et RODI~GUEz-OUBINA subass. nova hoc loco (CASASECA 1959: 325, table 1: relevr 4; relev6 27 in table 7 of this article). Table 7: relev6s 18-27. Holotypus: the same as that of the association: table 1:relev6 4 (CASASECA 1959: 325)

polypodietosum interjectl ORTIZ et RODRIGUEZ-OUBINA subass, nova hoc lt)co (CASASECA 1959: 325, table 1: releves 1, 2, 3 and 5; relev6s 14-17 in table 7 of this article). Table 7: relev6s 1-17. Holotypus: table 1:relev6 1 (CASASECA 1959: 325)

The pteridophytic association Gymnogrammo-Davallietum canariensis is very common on artificial walls and moist crags which have a compact moss layer - dominated by the bryophytes Homalotecium sericeum HEDW., Tortula rnuralis HEDW., Bryum bicolor DICKS., Targionia hypophylla L., Porella platyphylla (L.) P~IFF., Lejeunea cavifolia (EHRH.) LI~DB., etc., which are abundant in Galicia due to the effects of the high humidity, especially in shaded areas.

Within this association, two subassociations can be clearly recognised, one of which is characteristic of sheltered areas of the Thermocolline subbelt, where the climate is mild and free of frost and has permitted the survival of the macaronesian relict taxon Davallia canariensis (L) SM., whose presence characterizes this subassociation, and the other widely distributed in the Eucolline and Upper colline subbelt as well as some areas of the thermocolline subbelt where the adverse climate has impeded survival of Dava/l/a canar/ens/s.

Of the 5 relev6s included in the original table, Davallia caaariensis (L,) SM. is only present in one (relev6 4). Since the name of the association is Uymnogrammo-Davallietum canariensis, the selection of relevr 4 as the typus of this association name and of the typical subassociation Gymnogrammo-Davallietum davallietosum canariensis appears to be the most coherent choice. For the other subassociation we have selected the name

ORTIZ AND RODR~GUI~-OUBI~ SYNOPSIS OFTHE RUPICOLOUS VEGL~ATION 41

Table 8. P ~ ca~_';~m community

Rele~ nmaber 1 2 3 4 5 X~ede ( - " . ) 50 50 50 30O 300 Orientation SW SW N W W ~ , t ~ (o) 9o 9o 9o 9o 9o A~.~ . ~ d (m 2) 1 1 2 1 1 Cover of vasce~ plants (%) 60 7O 30 25 30 Average height of vegetation (can) 10 15 15 10 10 Number of spedcs per relev~ 8 7 5 5 4

Diagllo6tir taxa of :dli~tncr and order

~ / d a n u s 2.1 2.2 + 1.1 2.1 posietariajudaica 1.1 1.1 1.1 + O)mba/a~ mura//s 1.1. + + 2.2

D i a g n o ~ taxa of class

~ t r / c ~

Other taxa

1.1

P ~ c a ~ t ~ , m 33 4.4 ~ 2.1 r ] ~ ~ . ~ 1.1 + OwJ/don/ummajus + 1.1 Gowdumrobert/anum + +

Ho/cus/anatus +

2.1 +

Gymnopwnmo-Daval//etum canadens/s po/ypod/etosum /merject/with the observation that the taxon which appears in the mentioned table as Polypodium vu/gare L., belongs to P. /nterjectum SarVAS.

Besides 5 relev6s published by BELLOT and ~ (1959, Tab. 1) we present 22 original relev~s of this association in Tab. 7, of which, numbers 1 to 13 correspond to the subassociation Gymnogrammo-Davallietum po/ypod/etoswn/merject/and numbers 18 to 27 to the subassociation Gymnoffwnnw-DavaUietum davallietoswn omaden~.

As we stated at the outset, this siliceous association occurs mostly on art i f ic ial walls, where it is richest in species (relev~s 1-9, 18-21), as well as on natural crags, where it is poorer in species (relev~s 10-11, 22-24), and, in epiphytic form, on tree trunks, with generally no more than 2 or 3 species per relev~ (relev~s 12-13, 2.5-26).

On the basis of our data, the existence of the association Silene acutifolia-Holcus gayanus BELLOT 1951, which BELLOT (1966) considered to be related to Gymnosrwnmo-Dava///etum canar/ens/s, is not sufficiently verified.

42 FOI2A GEOBOTANICA El" PHYTOTAXONOMICA, 28, 1993

Table 9. Rume~ som,_,,_,s subsp, ga//aec/cus community

Relev~ number Altitude (m s.m.) Orientation Inellnation (o)

Cover of vascular plants (%) Average height of vegetation (era)

Number of species per relev~

1 2 3 4 100 100 80 80 NE N N N

60 70 70 70 10 10 10 10 50 40 70 50 20 15 20 15

8 8 12 13

Diagnostic taxa of order and elate

Rumor so ; t~ ,c subsp, gallaecicus L o ~ Sareosus

Other species

Sedum angticum Anthy///s vu/nezm/a subsp./be.z/ca Koe/er/a ghmca ./as/one montana

Sonchus o/erace~ Daucua carota subsp, gummifer Hypocho~ radicata Dacty~ g~zata Holcus mollis Si~.ne uniflora Leontodon taraxacoides Festuca mbra subsp.pminosa Picris echioides Rumex acetosa subsp, bifformis Silene scabrifiora subsp, gz//aec/ca L/nat/a at~ttono~/s

3.2 2.1 1.1 1.1 1.1 1.2 1.2

2.3 +

+.2 +.2 +

2.1 2.3 3.3 2.2 2.3 2.1 1.1 2.1 1.1

I . i +.2 + + + + + +

+

1.1 + 1.1 + I.I + + +

+ -l-

+ +

1.2

Polygonuln capitatu~ community Table 8

This is a community which occurs on artificial walls, in urban habitats of the Portuguese area of the Mineasian subsector (Galaico-Portuguese sector) where its optimum area of development is found. It is characterized by the abundant presence of the adventive species from the Himalayas Polygonum cap/tatum D. DoN. The rest of the floristlc composition does not differ very much from that ofParietarietum judaicae ~ 1928.

We have found this community only in the extreme southwestern area of Galicia.

ORTIZ AND RODRtGUEZ-OUBINA: SYNOPSIS OF THE RUPICOLOUS VEGETATION 43

Table 10. Rum~ suffmt/cosus community

Number of species 28 Number of relev~ 4

Diagnostic taxa of :~lllance

/~mex suffna/cosus 4 Phalacrocarpum oppositifolium 4

Diagnostic taxa of order and class

~ or/sin 1 Paronychia polygonifolia 1

O t h e r taxa

Deschampsia flemosa 2 Festuca indigesta s. L 2 Oi#taism,m,r.a 2 Coincya p s ~ subsp, set/gem 2 Linaria saxatilis 2

Taxa present in only one relev~: Anarrhinum beUidifolium, Calluna vulgaris, Crepis larnpsanoides, Eryngium duriaei, Festuca cf. iberica, Gentiana lutea, lurinea humilis, Leontodon hispidus, IAtzula caespitosa, Sedum anglicum, Se~rn brevifolium, Sedum hirsutum, Sedum tenuifolium, Sempervivum cantabricum, Sesamoides minor, Senecio adonidifolius, Solidago virga- aurea, Thymelaea dendrobtyum, I~ola riviniana. Column: Izco et ORTtZ (1987: 61, table 2)

Rumex scutatus subsp, gallaecicus community Table 9

This community colonises the serpentine dunite and eclogite mobile rock slides on the cliffs of the A Capelada peninsula, in the north coast of Galicia. It is characterized by the presence of Rumex scutatus L. subsp, gallaecicus LAGO, which is endemic to this locality.

Its syntaxonomic rank and position are problematic, among other reasons, because the independence of the serpentinicolous populations of Rumex scutatus at a subspecific level is not clear (L6PEZ Gor~zAtEZ 1990). On the other hand, plants belonging to Thlaspietea rotundifolii BR.-Bt. 1948 and its subdivisions are scarce in floristic composition of the community.

44 FOI..IA GEOBOTANICA L~r VHYTOTAXoNoMx'~, 28,1993

Table 11. Se.swnoido-Silenemm gayanae Izo3 ct ORTIZ corr. ORTIZ et RODRIGUEZ-OUBn~A

- subass, s//enetosum gayanae IZCO et ORTIZ in ORTIZ ct RODIdGUEZ-OUBIKIA (columns A, B) - sld)a~. ~ ~ ORTIZ ct RODRIGUI~-OuBI~ (relcv6s 1, 2)

Rclev6 number A B 1 2 Altitude (m s.m.) 1700 1750 Orien ta t ion W W

[Delln~tiOll (O) 45 50 Area Jmab~ (m2) 20 Cover of vascular plants (%) 10 20 Average height ot vegetation (an) 5 5 Set of relev~ 10 3 - Number of speci~ per reaev6 9 10

Diagnostic tam of association and mni~ncc

S//ene f o e t ~ subsp, gayana V 3 Sesamo/d~ minor V R u m a s u ~ u ~ 1I 1

Diagnostic tam of ordcr and class

L/naSa a/p/ha lI 2

DifferenfJ! taxa of subass. ~

Let~c'~m ~ subsp, r~erchon//

Other taxa

Su,n~ te,,.Co~us Ag~xttis uunc~_,_~_ subsp. ~ta.iaa

L e u c ~ p a U ~ subsp.j~.~,~ Sedum anglicum subsp.pyrena/cum Festuca bsdige.rm s.a. ~ o , uoaon ~ sub~p, c.,s:~,Vcus

Teesdaliopsis conferta 1as/one o/spa subsp, brev/sep~a

Desc~J~s/a fle~osa

V 2 IV 2 HI 3 IlI 3 HI 2 m

I 2 lI I 3 I 2

H II II II

LI 1.2

+ +

1.1 L1

1.2 +

+ 1.1

+

+

+

ORTIZ AND RODRIGUEZ-OUBI/7,1A: SYNOPSIS OF THE RUPICOLOUS VEGETATION 45

Relev6 number A B 1 2

Agrost/s t//en/ Jas/one/aev/s subsp, car/~tana Sot/dago v/~-aw~ D/anthus/angeanus Ca~ea,,uta h~an~/ Murbeckiella bo~ Hemiaria scabrida 1.1 +

Taxa present in only one relev6: ASrosas sto/on/fera+ .2, in Column A. Anarrh/num bellidifolium +, in Cohlmn A. Ou~enorrh/num m/nus 1.1, in Relev6 2. Ojffsus oromediterraneus +, in Column A. Dryopteris oreades +, in Column A. Festuca iberica +, in Relev6 1. Galium harcynicum subsp, vivianum r, in Column B. Hieracium pilosella + .2, in Column B. Linwia sup/ha +, in Relev6 1. Lotus comiculatus +.2, in Column B. Luzula caespitosa 1.3, in Column A. Nardus sO/cta r, in Column B. Plantago alpina war. eriopoda +, in Column B. Poa pratensis + .2, in Relev~ 1. Se&an album subsp. m/cranthum +, in Column A. Spcgula morisonii +, in Column A. Thyme/aea dendrobryum +.2, in Colnmn B.

Column. A: Izco et ORTIZ (1987: 62, table 3) Column. B: ORTtZ (1986: 138, table 24) Relev6s I and 2: Macizo de Pena Trevinca, Carballeda (Ourense). I.YrM: 29TPG7592. On artificial clay schists.

15. Cryptogrammo-Silenetum gayanae FERNANDEZ PRIETO COLT. ORTIZ et RODRIGUEZ-OUBINA hoe loco (FERNANDEZ PRETO 1983: 499, table 7)

We hereby correct the name of the association Cryptogrammo-Silenetum herminii FE~Ar~EZ PRIMO 1983, in accordance with Article 43 of the Code of the Phytosociological Nomenclature, since the taxon which forms part of the name of this association does not correspond to the Sitenefoetida L I ~ type (with which S. herm/n/is synonymous) but to S//ene foet/da subsp, gayana TALAV'EgA (TALAVERA 1988, 1990)

Rumex suffnaicosus community Table 10

This community is flofistically characterized by Rumex suffruticosus GAY ex MEISSNER and Phalacrocarpum oppositifolium (BRoT.) Wn.LX. subsp, oppositifolium. It occurs in the Oromediterranean belt principally over dry mobile rock slides, principally quartzitic, formed of medium-sized rock fragments. Although initially Izco et ORrtZ (1987) included it into the association Phalacrocarpo-Rumicetum suffruticosi RIVAS-MARTINEZ 1981, we now think that this latter is restricted to the Serra da Estrela (Portugal) where it is distinguished by the endemic plants Silenefoetida LINK subsp.foet/da and Senecio cespitosus BROT.

46 FOtJA OeO~OTA~CA ET pttYrorAXONOMICA, 28, 1993

16. S e s a m o i d ~ S ~ u n gayanae I ~ o et OsT~z corr. OSTtZ et RODR~GUEZ-OU~ hoc 1OCO (Izco et ORTIZ 1987: 62, table 3). Table 11.

silenetosum gayanae Izco et ORTIZ in OSTXZ et R O D ~ O U E Z ~ subass, nova hoc loco (Izco et OSTIZ 1987: 62, table 3: relevCs I-I0). Table 11: synoptic colnmn~ 1-2, Holotypus: the same as that of the association: table 3: relev~ 8 (Izc~ et Os~.~ 1987: 62)

lepidietosum reverchonii OR~Z et RODPJGUEZ-OUBI~ subass, m v a hoc loco Table 11: relev6s 1-2 Holotypus: table 11: relev~ 2

For the same reason as above (15), we hereby correct the association name Sesamoido-Silenetum henninii Izco et ORTIZ 1987.

The subassociation Seswnoido-Sgenetwn hermb~ii lepidietoswn revewJu~ whose differential taxon is Lepid/mn villarsii GREN. et GODSON subsp, revochoni/ (DEBEAUX) B m s ~ . , colonizes artificial roadside mobile rock slides made up of small frasments of slate covered w/th soil, and therefore, more nitrified in contrast to the behav/our of the subassodation type that occurs on similar banks, but of a natural origin and not or little nitrified.

We have observed the new subassociations in the Macizo de Penn Trevinca, at lower levels of the Oromediterranean bioclimafic belt, in the potential area of the Genisto-lw6peretwn nanae FERSA~3EZ PRr~O I983.

ACKNOWI .)~r)GEMENI~

The authors arc very gratef~ to Dr. lace for ~ collaboration in the elaboration of this work, to Dr. Am~o for his help ia designing the bioclimatic map, and to Dr. Juan Reinoso for identification of the bryophytes.

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ORTIZ AND RODR~GUI~-OuBI~: SYNOPSIS OFTHE RUPICOLOUS VEGETATION 47

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TLrI'IN T.G. et al. (1964-1980) [e, ds.]: Flora Europaea vol. 1-5. - Cambridge.

Re~iveA 14 January 1992, accepted 22 October 1992

48 FOLIA GEOmYrANICA El" PHYTOTAXONOMICA, 28, 1993

Appendix

Localisafion of relev~ and their geological substratum

Table 4

1. Muxfa (A Corufia). UTM: 29TMH8171. Muscovite and biotite granites. 2. II/as,Cies, V'q~o (Pontevedra). UTM: 29TNG0367. Muscovite and biotite granites. 3. Ponteceso (A Coru~). UTM: 29TNH0586. Granodiorites. 4. llla~ Cies, Vigo (Pontevedra). UTM: 29TNG0367. Granodiorites. 5. A l_qnT,~da, O Grove (Pontevedra). UTM: 29TNH0300. Muscovite and biotite granites. 6. Nereid, Muxfa (A CoruiIa). UTM: 27rMH7861. Metasediments.

Table 5

Relev6.s I and 2 top of Pana Trevinca Sur, A Veiga (Ourense). UTM: 29TPG8178. OrthoquartT/tes. Relev~s 3, 4, 8 and 9. Pena Trevinca Norte, Carballeda (Ourense). UTM: 29TPG8280. Orthoquartzites. Relev~ 5-7 ( ~ E 2 PRIETO 1983, table 5: relev~s 1-3). Relev~ 10. Top of Pico Maluro, Carbaneda (Ourense). UTM: 29TPG7883. Orthoquart2ites

Table 6

1. Embalse de Portodemouro~ Arz~ (A Coru~). UTM: 29TMH6846. Biotite schists. 2. Crecente (Pontevedra). UTM: 29TNG6366. Biotite granites. 3. Cudeiro (Ourense). UTM: 29 G9492. Biotite granites. 4. Ribadavia (Ourense). UTM: 29TMG708I. Biotite granites. 5. Embalse do Eume, As Pontes de Garcfa Rodrfgucz (A Coru~). UTM: 29TNJS006. Phyilites. 6. Close to Baamonde, Bogonte (Lugo). UTM: 29TPH008L Phyilites. 7. Pazos, Silleda (Pontevedra). UTM: 29TNH5384. Metamorphic dayey schists. 8. Berredo, A Golada (Pontevedra). UTM: 29TNH7942. Metamorphic clayey schists. 9. Rebordelo, Valpaco (Bra~anca, Portugal). UTM: 29TPG5124. Phyilkes. 10. Ourease. UTM: 29TNG9287. Metamorphic clayey schists. 1L Belesar, Chantada (Lugo). UTM: 29TPH0518. Diorites. 12. Salvaterra do Miio 0Pontevedra). UTM: 29TNG4159. Biotite granites. 13. Sta. Maria, RfDadavia (Ourense). UTM: 29TNG738Z Metamorphic clayey schists. 14. Embalse dos Peares, Carballedo (I.ego). UTM: 29TPH0509. Metamorphic clayey schists. 15. Close to Carballe~ Pifior (Ourense). UTM: 29TNHS108. Metamorphic clayey schists.

Table 7

1. Pontedeume (A Coru~). L �9 M: 29TNJ6806. Biotite schists. 2. Curtis (A Corufia). UTM: 29TNH6974. Muscovite and biotite granites. 3. Negreira (A Coru~). UTM: 29TNH2150. Glandular gneissic granites. 4. Noia (A Condta). UTM: 29TNH0937. Glandular biotite granites. 5. Muros (A Coru~). UTM: 29TMH9535. Orientated muscovite and biotite granites. 6. Santiago de Compostela (A Corufia). UTM: 29TNH3647. Glandular gneissic granites.

ORTIZ AND RODP.JGUEZ-OUBI~ SYNOPSIS OFTHE RUPICOLOUS VEGETATION 49

7. Santiago de Compostela (A Corufia). UTM: 29TNH3647. Muscovite and biotite granites. 8. Santiago de Compostela (A Coru~). UTM: 29TNH3647. Muscovite and biotite granites. 9. Vilaxoan, Vila~arcfa de Arousa (Pontevedra). UTM: 29 TNH1715. Muscovite and biotite granites. 10. l~,,tano, Portas (Pontevedra). UTM: 29TNH2613. Biotite granites 11. Ponteareas (Pontevedra). UTM: 29TNG4169. Biotite granites. 12. Santiago de Compostela (A Contea). UTM: 29TNH3647. On oak trunk. 13. Zaman~ Vi8o 0Pontevedra). LrFM: 29TNG2567. Glandular gneisses. 14-17. (BEI.Ixyr et ~ 1959, table 1: relev6s 1, 2, 3 and 5). 18. San Andr6s de Teixido, Cedeira (A Corufia). UTM: 29TNJ8240. Granulites. 19. Eirf~ Pontedeume (A Corufia). UTM: 29TNJ6908. Glandular gneissic granites. 20. Sobradelo, V'dagarcfa de Arousa (Pontevedra). UTM: 2ffI'NH1815. Metamorphic clayey schists. 21. Cambados (Pontevedra). UTM: 29TNH1507. Biotite granites. 22. Souto de Polo, Lantaiio, Portas (Pontevedra). UTM: 29TNH2613. Biotite granites. 23. Mte. Castro, Portas (Pontevedra). UTM: 29TNH2714. Biotite granites. 24. Mte. Lobeira, V'dagarcfa de Arousa (Pontevedra). UTM: 29TNH2012. Biotite granites. 25. Mte. Meda, Padr6n (A Corufia). UTM: 29TNH3033. On oak trunk. 26. Figueiras, Santiago de Compostela (A Corufia). UTM: 29TNH3450. On oak trunk. 27. ( B ~ et C.ASASEC~ 1959, table 1:relev6 4)

Table 8

1. Tabag6n, A Guarda (Pontevedra). UTM: 29TNG1817. Muscovite and biotite granites. 2. Tabag6n, A Guarda (Pontevedra). UTM: 29TNG1817. Muscovite and biotite granites. 3. V'danova da Cerveira (Minho, Portugal). UTM: 29TNG2143. Muscovite and biotite granites. 4. Braga (Minho, Potugal). UTM: 29TNG5200. Granodiorites. 5. Braga (Minho, Portugal). UTM: 29TNG5200. Granodiorites.

Table 9

1-4. Cabo Ortegal, Cariiio (A Coru~). UTM: 29TNJg148. Eclogites and Dtmites.